Kyle Miller obtained his PhD from the University of College London, working in the lab of Julie Cooper at the London Research Institute. His PhD work focused on studying telomeres and their roles in replication and genome maintenance in fission yeast. He then was a postdoctoral fellow at the University of California San Francisco focusing on chromatin and genome stability in budding yeast in the lab of Dr. David Tocyzski. He next completed another postdoctoral training in the lab of Prof. Stephen Jackson at the Gurdon Institute, University of Cambridge, England. This work focused on understanding the role of histone modifications and chromatin modifying enzymes in DNA double-strand break repair.