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1985: DSc, University of Newcastle-upon-Tyne。 1979: MA, University of Oxford。 1975: PhD University of Newcastle-upon-Tyne。 1973: MD, University of Newcastle-upon-Tyne。 1965: MB BS, Durham University。
2012-Present: Professor of Histopathology, Barts and the London School of Medicine and Dentistry, Queen Mary University of London。 1988–2011: Group Leader, Histopathology Unit, Cancer Research UK London Research Institute, UK。 2001-2011: Warden, Barts and the London, School of Medicine and Dentistry, Queen Mary University of London UK。 2001–2011: Professor of Investigative Science, Barts and the London School of Medicine and Dentistry, Queen Mary University of London UK; Honorary Consultant Histopathologist, Barts and the London NHS Trust; UK 。Visiting Chairs at University College London and University of Oxford。 1997–2000: Deputy Principal and Vice-Principal for Research, Imperial College School of Medicine, UK。 1996–1997: Dean, Royal Postgraduate Medical School, UK。 1988-1996: Director of Clinical Research, ICRF, UK。 1980–1996: Professor of Histopathology, Royal Postgraduate Medical School, UK; Director of Histopathology, Hammersmith and Queen Charlotte's SHA, UK; Honorary Consultant Histopathologist, Hammersmith and Queen Charlotte's SHA, UK。 1997-1979: Nuffield Reader in Pathology, University of Oxford, UK; Honorary Consultant Pathologist, Oxford University Hospital Group, UK; Fellow, Green College, UK。 1975-1976: Senior Lecturer in Pathology, University of Newcastle-upon Tyne, UK; Honorary Consultant Pathologist, Newcastle University Hospitals Group, UK。 1971-1975: Lecturer in Pathology, University of Newcastle-upon-Tyne, UK。
publication:
Gaisa NT, Graham TA, McDonald SA, Cañadillas-Lopez S, Poulsom R, Heidenreich A, Jakse G, Tadrous PJ, Knuechel R, Wright NA. The human urothelium consists of multiple clonal units, each maintained by a stem cell. J Pathol. 2011; PMID: 21744343
Graham TA, Humphries A, Sanders T, Rodriguez-Justo M, Tadrous PJ, Preston SL, Novelli MR, Leedham SJ, McDonald SA, Wright NA. Use of methylation patterns to determine expansion of stem cell clones in human colon tissue. Gastroenterology. 2011; PMID: 21192938
Leedham SJ, Graham TA, Oukrif D, McDonald SA, Rodriguez-Justo M, Harrison RF, Shepherd NA, Novelli MR, Jankowski JA, Wright NA. Clonality, founder mutations, and field cancerization in human ulcerative colitis-associated neoplasia. Gastroenterology. 2009; PMID: 19103203
Leedham SJ, Preston SL, McDonald SA, Elia G, Bhandari P, Poller D, Harrison R, Novelli MR, Jankowski JA, Wright NA. Individual crypt genetic heterogeneity and the origin of metaplastic glandular epithelium in human Barrett's oesophagus. Gut. 2008; PMID: 18305067
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