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From then onwards, he collaborated closely with David Warrell in Nigeria, Thailand, Sri Lanka, Papua New Guinea, Ecuador, Peru and, notably, during an EEC-funded programme with Instituto Butantan, in Brazil from 1989 until 2000. He pioneered EIA for detection and quantitation of venom antigens in clinical samples, established a panel of WHO standard tests to characterise medically-important venoms and antivenoms and has pursued innovation in antivenom design, production and testing. He has provided in-the-field and laboratory support for clinical studies of snake-bite and arthropod bites and stings in many tropical countries. At LSTM he encouraged the introduction of molecular techniques for the characterisation of important snake-venom components identified by his group. He was a key element of a sustained and ultimately successful collaboration between LSTM, the University of Oxford and the Federal Ministry of Health, Nigeria, the “EchiTAb” project, to develop new antivenoms for treatment of Echis ocellatus envenoming. Throughout his career he has proved an inspiring teacher, trainer, research supervisor, consultant and collaborator, while his warm Yorkshire personality and sense of humour has endeared him to his many international friends and colleagues.
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