Bifurcation Analysis of an Ecological System with State-Dependent Feedback Control and Periodic Forcing

By assuming a periodic variation in the intrinsic growth rate of the prey, a nonlinear ecological system with periodic forcing and state-dependent feedback control is proposed. The main purpose of the present paper is to study the dynamical behavior generated by periodic forcing and nonlinear impulse perturbations and their effects on pest control. To do this, we first investigate the existence and stability of the boundary periodic solution, and then we employ the numerical bifurcation techniques, mainly including one-dimensional and two-dimensional parameter bifurcation analyses, to reveal that the system exhibits rich and complex dynamic behaviors. Especially, period-adding bifurcation with chaos is found in the two-parameter bifurcation plane. Moreover, we find the periodic structure similar to Arnold tongues, and they are arranged according to the sequence of a Farey tree. In addition, one-dimensional bifurcation diagrams reveal the existence of order-k periodic, and chaotic solutions, multiple coexisting attractors, period-doubling bifurcations, period-halving bifurcations, and so on. Finally, the effects of the initial population density of pests and natural enemies on the pulse frequency and the biological significance related to the numerical results are studied and discussed.