Ichnological response to the Middle Eocene Climatic Optimum (MECO) in the Bartonian deposits of Kutch Basin, India

The Middle Eocene Climatic Optimum (MECO) represents a short-lived interval of global warming that disrupted the long-term cooling episode extending from the middle to late Eocene time. Biostratigraphically the MECO corresponds to the upper interval of planktic foraminiferal zones E11–E12 and the shallow benthic zone 17, both of are well-documented within the Bartonian mixed siliciclastic‑carbonate sequence of the Kutch Basin in western India. This case study aims to investigate the impact of MECO on the endobenthic community using ichnology as a proxy. The studied sequence comprises the underlying Harudi Formation, consisting of the marine green and grey shale and bioclastic limestone beds, and the overlying Fulra Limestone characterized by benthic foraminifera-rich limestones. The Harudi Formation is bereft of trace fossils except in one bioturbated firmground and two bioeroded hardground horizons. The MECO corresponds to the thin carbonate packstone, the middle green shale, and the Nummulites obtusus rudstone occurring at the middle to upper part of the Harudi succession established earlier by a concurrent negative δ13Corg isotopic excursion. The shelly limestone or bivalve coquina bed deposited immediately prior to the beginning of the MECO consists of a monospecific firmground ichnofabric characterized by Balanoglossites triadicus. Out of two bioeroded beds, the lower one coincides with the earliest MECO interval and is marked by a brown carbonate hardground colonized with two ichnospecies of Gastrochaenolites. The upper bioeroded horizon corresponding to the end of the MECO is characterized by a series of top-bioeroded (with Gastrochaenolites, Entobia, and Oichnus) carbonate nodules containing multigenerational septarian cracks. The upper hardground demarcates the top of the Nummulites obtusus coquina bed and the starkest basin-wide sediment starvation, making it a major condensed horizon that developed following a steady glacio-eustatic rise during the warming event. The MECO prompted the subsequent deposition of shallow-marine platform carbonates of the Fulra Limestone by accelerating a sharp rise in the sea-level and proliferation of diverse Larger Benthic Foraminifera in a warm oligotrophic seawater.