A Complex Patterned-Ground Ecosystem in the Sierra Nevada, California, USA Supports Unusual Habitat for American Pika

A complex landscape lies on an alpine plateau in the high Sierra Nevada, California, and comprises spatially organized physical and ecological features that interact to create a unique ecosystem. At the upslope end a tree-limited ribbon forest of whitebark pines grows on the crest of a short, steep slope, which is the only deformation on the otherwise flat plateau. A long-persistent snowbank forms on the slope in the lee of the ribbon forest; meltwater provides moisture to support a productive wet meadow just below. Below the meadow is a narrow, linear "rampart" comprising large sorted stones, and below this lies a large patterned-ground area of cryogenically sorted circles. The soil domains are densely covered with species-rich vegetation and are abruptly segregated from stone domains. The latter serve as distributed springheads in the otherwise dry patterned-ground terrain. American pikas have colonized the rocky nets of the stone domains and forage plants of the soil circles. The unusual context of this landscape enables pika territories to be more tightly packed than in traditional habitat. We propose a scenario for the Pleistocene origin of this ecosystem and a unique lithologic subsurface barrier that contributes to spring formation during warm intervals.