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The Rockefeller University Press $30.00 J. Cell Biol. Vol. 209 No. 2 289–303 www.jcb.org/cgi/doi/10.1083/jcb.201409032 JCB 289 Correspondence to Dana T. Graves: dtgraves@dental.upenn.edu Abbreviations used in this paper: AGE, advanced glycation end product; CCL20, chemokine (C-C motif) ligand 20; ChIP, chromatin immunoprecipitation; CML, carboxymethyllysine; FOXO1, forkhead box O1; NHEK, normal human epidermal keratinocyte; PCNA, proliferating cell nuclear antigen; SERPINB2, serpin peptidase inhibitor, clade B (ovalbumin), member 2; uPAR, urokinase-type plasminogen activator receptor. Introduction