Hepatic DPP 4 DNA-methylation associates with fatty liver Running title : Epigenetic regulation of hepatic DPP 4

* Both authors contributed equally to this work 1 German Institute of Human Nutrition Potsdam-Rehbruecke, Department of Experimental Diabetology, Nuthetal, Germany 2 German Center for Diabetes Research (DZD), München-Neuherberg, Germany 3 CNRS UMR 8199, Lille Pasteur Institute, Lille, France 4 Lille University, Lille, France 5 European Genome Institute for Diabetes, Lille, France 6 Department of Genomics of Common Disease, School of Public Health, Hammersmith Hospital, Imperial College London, London, U.K. 7 INSERM UMR 1190, CHU Lille, Lille, France 8 Epigenetics and Diabetes Unit, Department of Clinical Sciences, Lund University Diabetes Centre, Malmö, Sweden 9 Institute of Public Health and Clinical Nutrition, Clinical Nutrition, University of Eastern Finland, Kuopio, Finland 10 Clinical Nutrition and Obesity Center, Kuopio University Hospital, Kuopio, Finland