Indian genetic heritage in Southeast Asian populations 1 2

2 Piya Changmai, Kitipong Jaisamut, Jatupol Kampuansai, Wibhu Kutanan, N. Ezgi Altınışık, Olga 3 Flegontova, Angkhana Inta, Eren Yüncü, Worrawit Boonthai, Horolma Pamjav, David Reich, Pavel 4 Flegontov 5 6 7 1 Department of Biology and Ecology, Faculty of Science, University of Ostrava, Ostrava, Czech Republic 8 2 Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand 9 3 Research Center in Bioresources for Agriculture, Industry and Medicine, Chiang Mai University, Chiang Mai, 10 Thailand 11 4 Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, Thailand 12 5 Faculty of Sociology and Anthropology, Thammasat University, Pathum thani, Thailand 13 6 Hungarian Institute for Forensic Sciences, Institute of Forensic Genetics, Budapest, Hungary 14 7 Department of Human Evolutionary Biology, Harvard University, Cambridge, MA, USA 15 8 Department of Genetics, Harvard Medical School, Boston, MA, USA 16 9 Broad Institute of MIT and Harvard, Cambridge, MA, USA 17 Current Address: Human-G Laboratory, Department of Anthropology, Hacettepe University, Ankara, Turkey 18 19 * corresponding author, 20