A diverse Miocene fish assemblage (Chondrichthyes and Osteichthyes) from the Pécs-Danitzpuszta sand pit (Mecsek Mts, Hungary)

Chondrichthyans and osteichthyans are widely reported from marine sediments of the Central Paratethys, not only by sporadic occurrences, but also by complex, diverse fish assemblages. Here we present a rich fish fauna from the upper Miocene (Pannonian, Tortonian) lacustrine sediments exposed in the Pécs-Danitzpuszta sand pit, in the SW Pannonian Basin. Altogether 22 227 specimens were investigated, and they could be classified into 17 chondrichthyan and 16 osteichthyan taxa. Among the chondrichthyans Odontaspididae (55.51%) and Myliobatidae (14.4%) are the most abundant, while the bony fishes are dominated by the Sparidae (77.07%). The limonitic, yellow, coarse-grained, gravelly sands yielded an extremely large amount of isolated fish fossils. A considerable part of the vertebrate material of the sands is likely reworked from older, middle Miocene (Badenian and Sarmatian, i.e. Langhian and Serravallian) sediments. The late Miocene calcareous marls underlying the sands also yielded a variety of fish remains. From these remains, associated and articulated latid bones are coeval with the sediment and suggest freshwater and brackish conditions in the area, in accord with other biotic data. Other, isolated fossils, namely teeth, otoliths, cycloid scales and jaw elements of Gadidae, Gobiidae, Sparidae and Latidae could be reworked and thus have a very limited paleoecological significance. Based on habitat preferences of extant relatives, all chondrichthyans and most osteichthyan taxa found in the sands must be of Badenian origin. The abundant remains allow for some conclusions on the environmental conditions during the Badenian. They refer to a shallow, coastal environment with tropical-subtropical climate with connection to more open water habitats. Remains of some osteichthyan taxa were found in coprolites, showing that these taxa were part of the food chain as prey items. Sirenian ribs and odontocete limb bones bearing tooth marks refer to trophic relations between marine mammals and large sized macropredatory sharks. The Pécs-Danitzpuszta record of shark-attributed bite marks on bones of marine mammals is the second of the Badenian of the Central Paratethys. Acipenserids and latids of the sands, linked to brackish and/or freshwater environments, could originate from any Miocene stratigraphic units.