A look beyond the colour: taxonomic revision of Coilodes Westwood, 1846 (Coleoptera, Hybosoridae), with the description of six new species

Coilodes Westwood, 1846 is a Neotropical genus of Hybosorinae. Despite being a morphologically homogeneous genus, it presents a great deal of intraspecific variation, especially with reference to the colour of integuments. This makes identification of species even more difficult, since the majority of original descriptions present the colour as the main diagnostic character. To solve this problem, the first taxonomic revision of Coilodes is presented. The genus now comprises 13 species. Redescriptions of C. castaneus Westwood, 1846, C. fumipennis Arrow, 1909, C. humeralis (Mannerheim, 1829), C. niger (Mannerheim, 1829), C. ovalis Robinson, 1948, C. parvulus Westwood, 1846, and C. punctipennis Arrow, 1909 are presented. Coilodes niger (Mannerheim, 1829) has its status revalidated and three new synonyms are proposed: C. gibbus (Perty, 1830) and C. chilensis Westwood, 1846 with C. humeralis, and C. nigripennis Arrow, 1903 with C. castaneus. Biological and geographical distribution data are expanded. Lectotypes are designated for C. humeralis and C. niger. Six new species are described: C. bezerrai Basilio & Vaz-de-Mello sp. nov., C. edeiltae Basilio & Vaz-de-Mello sp. nov., C. lunae Basilio & Vaz-de-Mello sp. nov., C. mayae Basilio & Vaz-de-Mello sp. nov., C. ravii Basilio & Vaz-de-Mello sp. nov., and C. skelleyi Basilio & Vaz-de-Mello sp. nov. An identification key for the males of the species is presented.