This revision is part of a continuing series of taxonomic work aimed at the description of new taxa and the redescription of known taxa of the Tachinidae of Area de Conservación Guanacaste in northwestern Costa Rica. Here we describe 33 new species in the genus Belvosia Robineau-Desvoidy, 1830 (Diptera: Tachinidae). All species described here were reared from this ongoing inventory of wild-caught caterpillars spanning a variety of families (Lepidoptera: Erebidae, Eupterotidae, Noctuidae, Notodontidae, Saturniidae, and Sphingidae). We provide a morphological description of each species with limited information on life history, molecular data, and photographic documentation. In addition to the new species, the authors provide a redescription of the genus Belvosia , as well as provide a key to the identification of the species present in the Meso- and North-American fauna. The following 33 new species of Belvosia Robineau-Desvoidy, 1830, all authored by Fleming & Woodley, are described: Belvosia adrianguadamuzi Fleming & Woodley sp. n. , Belvosia anacarballoae Fleming & Woodley sp. n. , Belvosia angelhernandezi Fleming & Woodley sp. n. , Belvosia brigittevilchezae Fleming & Woodley sp. n. , Belvosia alixtomoragai Fleming & Woodley sp. n. , Belvosia carolinacanoae Fleming & Woodley sp. n. , Belvosia ciriloumanai Fleming & Woodley sp. n. , Belvosia diniamartinezae Fleming & Woodley sp. n. , Belvosia duniagarciae Fleming & Woodley sp. n. , Belvosia duvalierbricenoi Fleming & Woodley sp. n. , Belvosia eldaarayae Fleming & Woodley sp. n. , Belvosia eliethcantillanoae Fleming & Woodley sp. n. , Belvosia freddyquesadai Fleming & Woodley sp. n. , Belvosia gloriasihezarae Fleming & Woodley sp. n. , Belvosia guillermopereirai Fleming & Woodley sp. n. , Belvosia harryramirezi Fleming & Woodley sp. n. , Belvosia hazelcambroneroae Fleming & Woodley sp. n. , Belvosia jorgehernandezi Fleming & Woodley sp. n. , Belvosia josecortezi Fleming & Woodley sp. n. , Belvosia joseperezi Fleming & Woodley sp. n. , Belvosia keinoraragoni Fleming & Woodley sp. n. , Belvosia luciariosae Fleming & Woodley sp. n. , Belvosia manuelpereirai Fleming & Woodley sp. n. , Belvosia manuelriosi Fleming & Woodley sp. n. , Belvosia minorcarmonai Fleming & Woodley sp. n. , Belvosia osvaldoespinozai Fleming & Woodley sp. n. , Belvosia pabloumanai Fleming & Woodley sp. n. , Belvosia petronariosae Fleming & Woodley sp. n. , Belvosia ricardocaleroi Fleming & Woodley sp. n. , Belvosia robertoespinozai Fleming & Woodley sp. n. , Belvosia rostermoragai Fleming & Woodley sp. n. , Belvosia ruthfrancoae Fleming & Woodley sp. n. , Belvosia sergioriosi Fleming & Woodley sp. n. Belvosia canalis Aldrich, 1928 is reared and recorded from the inventory; new information relative to host is provided and the species is rediscribed. The following are proposed by Fleming & Woodley as new synonyms of Belvosia Robineau-Desvoidy, 1830: Brachybelvosia Townsend, 1927 syn. n. , Belvosiomimops Townsend, 1935 syn. n. The following three new combinations are proposed as a result of the new synonymies: Belvosia brasilensis (Townsend, 1927), comb. n. ; and Belvosia barbiellinii (Townsend, 1935), comb. n. The authors also propose the following new synonymies: Belvosia brasilensis (Townsend, 1927) = Belvosia aurulenta (Bigot, 1888), syn. n. ; Belvosia pollinosa Rowe, 1933 = Belvosia borealis Aldrich, 1928 syn. n. ; Belvosia weyenberghiana (Wulp, 1883) = Belvosia fuliginosa (Walker, 1853) syn. n. ; Belvosia brasiliensis Townsend, 1927 = Belvosia fuliginosa (Walker, 1853) syn. n. ; Belvosia luteola Coquillett, 1900 = Belvosia ochriventris (Wulp, 1890) syn. n. ; Belvosia socia (Walker, 1853) = Belvosia proxima (Walker, 1853) syn. n. ; Belvosia chrysopyga (Bigot, 1887) = Belvosia unifasciata (Robineau-Desvoidy, 1830) syn. n. ; Belvosia chrysopygata (Bigot, 1888) = Belvosia unifasciata (Robineau-Desvoidy, 1830) syn. n.

We revise the genus Metaplagia Coquillett, 1895 and describe five new species from Area de Conservación Guanacaste (ACG) in northwestern Costa Rica. All new species were reared from an ongoing inventory of wild-caught caterpillars spanning a variety of species within the family Sphingidae (Lepidoptera: Sphingidae). Our study provides a concise description of each new species using morphology, life history, molecular data and photographic documentation. In addition to the new species, the authors provide a re-description of the genus and a revised key to the species of Metaplagia . The following five new species of Metaplagia are described: Metaplagia leahdennisae Fleming & Wood sp. n. , Metaplagia lindarobinsonae Fleming & Wood sp. n. , Metaplagia paulinesaribasae Fleming & Wood sp. n. , Metaplagia robinsherwoodae Fleming & Wood sp. n. and Metaplagia svetlanakozikae Fleming & Wood sp. n. The following is proposed by Fleming & Wood as new combination of Plagiomima Brauer & Bergenstamm, 1891: Plagiomima latifrons (Reinhard, 1956) comb. n.

We describe 25 new species in the genusTelothyriavan der Wulp, 1890 from Area de Conservación Guanacaste (ACG) in northwestern Costa Rica. All species herein described were reared from an ongoing inventory of wild-caught caterpillars spanning two families (Lepidoptera: Crambidae, and Tortricidae). Our study provides a concise description of each new species using morphology, life history, molecular data, and photographic documentation; a redescription of the genus, and its type species as well as a revised key to species ofTelothyriaoccurring in the Mesoamerican region. We also suggest seven new synonymies resulting in 11 new combinations.The following 25 new species ofTelothyriaare described:T. aidanisp. n.,T. alexanderisp. n.,T. auranticrussp. n.,T. auriolussp. n.,T. bicuspidatasp. n.,T. carolinacanoaesp. n.,T. clavatasp. n.,T. cristatasp. n.,T. diniamartinezaesp. n.,T. duniagarciaesp. n.,T. duvalierbricenoisp. n.,T. eldaarayaesp. n.,T. erythropygasp. n.,T. fimbriatasp. n.,T. fulgidasp. n.,T. gloriashihezaraesp. n.,T. griseasp. n.,T. harryramirezisp. n.,T. incisasp. n.,T. manuelpereiraisp. n.,T. obscurasp. n.,T. omissasp. n.,T. osvaldoespinozaisp. n.,T. peltatasp. n., andT. ricardocaleroisp. n.The following are proposed by Fleming & Wood as new generic synonyms ofTelothyria:ComatactaCoquillettSyn. n.,FloradaliaThompsonSyn. n.,PtilomyiaCurranSyn. n.,PtilomyiopsisTownsendSyn. n.,PtilomyoidesCurranSyn. n.,EuptilomyiaSyn. n.,EutelothyriaTownsendSyn. n.The following new combinations are proposed as a result of the new synonymies:Telothyria bequaerti(Curran, 1925)Comb. n.,Telothyria cruenta(Giglio-Tos, 1893)Comb. n.,Telothyria frontalis(Townsend, 1939)Comb. n.,Telothyria insularis(Curran, 1927)Comb. n.,Telothyria itaquaquecetubae(Townsend, 1931)Comb. n.,Telothyria major(Thompson, 1963)Comb. n.,Telothyria micropalpus(Curran, 1925)Comb. n.,Telothyria minor(Thompson, 1963)Comb. n.,Telothyria nautlana(Townsend, 1908)Comb. n.,Telothyria plumata(Curran, 1925)Comb. n.,Telothyria trinitatis(Thompson, 1963)Comb. n.,Telothyria variegata(Fabricius, 1805)Comb. n.Musca tricinctaFabricius is synonymized underTelothyria variegataFabricius,Syn. n.Telothyria schineriFleming & Woodnom. n.is proposed as a replacement name forMiltogramma brevipennisSchiner.Additionally we provide redescriptions of two previously named species: the type speciesTelothyria cupreiventris(van der Wulp) due to its being the type species, andTelothyria relicta(van der Wulp) due to its having been reared as an outcome of the inventory.

We describe one new genus and its one new species from Area de Conservación Guanacaste (ACG) in northwestern Costa Rica. Our study provides a concise description of this new species using morphology, life history, molecular data and photographic documentation. Chorotegamyiagen. n. is described, along with its type species, Chorotegamyia aureofacies sp. n. A modified key to the Uramyini is given to further elucidate the tribe.

We describe 22 new species in the genus Hyphantrophaga Townsend, 1892 (Diptera: Tachinidae) from Area de Conservación Guanacaste (ACG) in north-western Costa Rica. All species were reared from an ongoing inventory of wild-caught caterpillars spanning a variety of families (Lepidoptera: Bombycidae, Crambidae, Depressariidae, Doidae, Erebidae, Euteliidae, Gelechiidae, Geometridae, Hedylidae, Hesperiidae, Immidae, Lasiocampidae, Limacodidae, Megalopygidae, Mimaloniidae, Noctuidae, Nolidae, Notodontidae, Nymphalidae, Papilionidae, Pieridae, Phiditiidae, Pterophoridae, Pyralidae, Riodinidae, Saturniidae, Sphingidae, Thyrididae, Tortricidae and Zygaenidae). We provide a morphological description of each species together with information on life history, molecular data and photographic documentation. In addition to the new species, we provide a redescription of the genus, as well as the redescription of three previously described species, which were also collected within ACG during this study: Hyphantrophagaangustata (van der Wulp), Hyphantrophagamyersi (Aldrich) and Hyphantrophagavirilis (Aldrich & Webber). The following 22 new species of Hyphantrophaga are described: Hyphantrophagaadrianguadamuzi Fleming & Wood sp. n., Hyphantrophagaalbopilosa Fleming & Wood sp. n., Hyphantrophagaanacordobae Fleming & Wood sp. n., Hyphantrophagacalixtomoragai Fleming & Wood sp. n., Hyphantrophagacalva Fleming & Wood sp. n.., Hyphantrophagaciriloumanai Fleming & Wood sp. n., Hyphantrophagadanausophaga Fleming & Wood sp. n., Hyphantrophagadiniamartinezae Fleming & Wood sp. n., Hyphantrophagaduniagarciae Fleming & Wood sp. n., Hyphantrophagaedwinapui Fleming & Wood sp. n., Hyphantrophagaeldaarayae Fleming & Wood sp. n., Hyphantrophagaeliethcantillanoe Fleming & Wood sp. n., Hyphantrophagagilberthampiei Fleming & Wood sp. n., Hyphantrophagaguillermopereirai Fleming & Wood sp. n., Hyphantrophagahazelcambroneroae Fleming & Wood sp. n., Hyphantrophagaluciariosae Fleming & Wood sp. n., Hyphantrophagamanuelriosi Fleming & Wood sp. n., Hyphantrophagamorphophaga Fleming & Wood sp. n., Hyphantrophaganigricauda Fleming & Wood sp. n., Hyphantrophagaosvaldoespinozai Fleming & Wood sp. n., Hyphantrophagapabloumanai Fleming & Wood sp. n. and Hyphantrophagasimilis Fleming & Wood sp. n. The following are proposed by Wood as new synonyms of Hyphantrophaga Townsend, 1892: Brachymasicera Townsend, 1911 syn. n., Ommasicera Townsend, 1911 syn. n., Ophirosturmia Townsend, 1911 syn. n., Patillalia Curran, 1934 syn. n. and Ypophaemyiops Townsend, 1935 syn. n. The following nine new combinations are proposed as a result of the new synonymies: Hyphantrophagaadamsoni (Thompson, 1963), comb. n., Hyphantrophagafasciata (Curran, 1934), comb. n., Hyphantrophagaglauca (Giglio-Tos, 1893), comb. n., Hyphantrophagagowdeyi (Curran, 1926), comb. n., Hyphantrophagamyersi (Aldrich, 1933), comb. n., Hyphantrophaganigripes (Townsend, 1928), comb. n., Hyphantrophagaoptica (Schiner, 1868), comb. n., Hyphantrophagapolita (Townsend, 1911), comb. n., Hyphantrophagasubpolita (Townsend, 1912), comb. n.

Twenty-three new species of the genus Calolydella Townsend, 1927 (Diptera: Tachinidae) are described, all reared from multiple species of wild-caught caterpillars across a wide variety of families (Lepidoptera: Crambidae; Erebidae; Geometridae; Hesperiidae; Lycaenidae; Nymphalidae; Pieridae; Riodinidae; and Sphingidae). All caterpillars were collected within Area de Conservación Guanacaste (ACG), in northwestern Costa Rica. This study provides a concise description of each new species using morphology, life history, molecular data, and photographic documentation. In addition to the new species, we also provide a generic redescription and revised key to species of the genus Calolydella from Central and South America.The following 23 new species of Calolydella are described by Fleming and Wood: C. adelinamoralesaesp. n., C. alexanderjamesisp. n., C. argenteasp. n., C. aureofaciessp. n., C. bicolorsp. n., C. bifissussp. n., C. crocatasp. n., C. destitutasp. n., C. discalissp. n., C. erasmocoronadoisp. n., C. felipechavarriaisp. n., C. fredriksjobergisp. n., C. inflatipalpissp. n., C. interruptasp. n., C. nigripalpissp. n., C. omissasp. n., C. ordinalissp. n., C. renemalaiseisp. n., C. susanaroibasaesp. n., C. tanyadapkeyaesp. n., C. tenebrosasp. n., C. timjamesisp. n., C. virginiajamesaesp. n.Lydella frugale Curran, 1934 is proposed as a new synonym of Pygophorinia peruviana Townsend, 1927, syn. n., under the combination Calolydella frugale (Curran, 1934), comb. n.

We describe five new species in the genus Vibrissina Rondani from Area de Conservación Guanacaste (ACG). All species were reared from wild-caught sawfly larvae (Hymenoptera: Symphyta: Argidae and Tenthredinidae). We provide a morphological description of each species together with information on life history, molecular data, and photographic documentation. Five new species of Vibrissina Rondani: Vibrissina randycurtisi sp. n., V. randyjonesi sp. n., V. robertwellsi sp. n., V. danmartini sp. n., V. hallwachsorum sp. n.

Abstract Background We describe a new species in the genus Voria Robineau-Desvoidy, 1830 (Diptera: Tachinidae: Voriini) from Area de Conservación Guanacaste (ACG) in northwestern Costa Rica. It was reared as part of an ongoing inventory of wild-caught caterpillars spanning a variety of moth and butterfly families (Lepidoptera). Our study provides a concise description of the new species using morphology, life history, molecular data, and photographic documentation. In addition to the new species, we provide a diagnosis of the genus as well as new data relating to host use. New information The following new species of Voria is described: Voria erasmocoronadoi Fleming & Wood sp. n. The following are proposed by Fleming & Wood as new synonyms of Voria: Xenoplagia Townsend, 1914 syn. n., Hystricovoria Townsend, 1928 syn. n., Afrovoria Curran, 1938 syn. n., and Anavoria Mesnil, 1953 syn. n., and Itavoria Townsend, 1931 syn. n. The following new combinations are proposed as a result of the new synonymies: Voria bakeri (Townsend, 1928), comb. n. and Voria setosa (Townsend, 1914), comb. n.The authors also propose Voria pollyclari (Rocha-e-Silva, Lopes & Della Lucia, 1999), comb. n. based on the morphology of the holotype.

BackgroundWe describe two new species in the genus Erythromelana Townsend, 1919 from Area de Conservacion Guanacaste (ACG) in northwestern Costa Rica. Both species were reared from wild-caughtcaterpillars of Eois spp. (Lepidoptera: Geometridae). We provide a concise description of each species using morphology, life history, molecular data, and photographic documentation.New informationErythromelana jimmychevezi Fleming & Wood sp. nov.

We describe a new species of Phosocephala Townsend, and provide a new collection record, and description of the previously unknown male, of Phosocephala metallica Townsend, from Area de Conservación Guanacaste (ACG), northwestern Costa Rica. All ACG specimens were reared from wild-caught Lepidoptera larvae (Lepidoptera: Erebidae, Nolidae). We provide a concise description of both species using morphology, life history, molecular data, and photographic documentation. The new species is authored and described by Fleming and Wood. Phosocephala alexanderi sp. n.

We describe three new species in the genus Ametadoria Townsend from Area de Conservación Guanacaste (ACG), Costa Rica. All three were reared from wild-caught Zygaenidae and Lacturidae caterpillars. We provide a concise description of each species using morphology, life history and molecular data, with photographic documentation. The new species are authored and described by Fleming and Wood: Ametadoria karolramosae sp. nov., Ametadoria leticiamartinezae sp. nov., and Ametadoria mauriciogurdiani sp. nov. The following are proposed by Wood as new synonyms of Ametadoria Townsend: Adidyma Townsend syn. nov., and Abolodoria Townsend syn. nov. The following new combinations occur as a result of these new synonymies: Ametadoria abdominalis (Townsend) comb. nov., Ametadoria austrina (Coquillett) comb. nov., Ametadoria humilis (Wulp) comb. nov., Ametadoria misella (Wulp) comb. nov. Ametadoria adversa (Townsend) is proposed as a junior synonym of ​Ametadoria unispinosa Townsend, syn. nov​.

We describe three new species of Trigonospila Pokorny (Tachinidae: Blondeliini) from Area de Conservación Guanacaste (ACG), northwestern Costa Rica. All were reared from -various species of ACG caterpillars during an ongoing inventory of caterpillars, their food plants and their parasitoids in dry forest, rain forest and cloud forest. By coupling morphology, photographic documentation, life history and molecular data, we provide a clear and concise description of each species. All species published as new, are known to be previously undescribed as a result of careful study of the genus by DMW. This study builds on the current knowledge of the genus by adding three new species to the current 7 described in the New World. Trigonospila edwinbermudezi sp. n., Trigonospila uniformis sp. n., and Trigonospila josemariamoragai sp. n. are all authored and described as new by Fleming and Wood, with a key to their identification. The authors also offer a new record and description of the previously unknown male of Trigonospila panamensis (Townsend), reared from ACG caterpillars.

Nine new species of Itaplectops Townsend (Diptera: Tachinidae) are described from Area de Conservación Guanacaste (ACG), northwestern Costa Rica. All specimens have been reared from ­various species of ACG caterpillars in the families Limacodidae and Dalceridae. By combining morphological, photographic, and genetic barcode data we provide clear yet concise descriptions. The following nine new species are described in the genus Itaplectops: Itaplectops akselpalolai, Itaplectops anikenpalolae, Itaplectops argentifrons, Itaplectops aurifrons, Itaplectops ericpalolai, Itaplectops griseobasis, Itaplectops omissus, Itaplectops shellymcsweeneyae, Itaplectops tristanpalolai. We move Itaplectops to the tribe Uramyini from its original placement within the Blondeliini, and we discuss its systematic placement. We also provide a key differentiating the, genera of the tribe Uramyini as well as the known species of Itaplectops.

We describe seven new species of Spathidexia (Diptera: Tachinidae) reared from Area de Conservación Guanacaste (ACG), northwestern Costa Rica. All were reared from -various species of ACG caterpillars during an ongoing inventory of caterpillars, their food plants and their parasitoids. By coupling morphology, photographic documentation, life history and molecular data, we provide a clear and concise description of each species. All are known to be previously undescribed as a result of a comprehensive study of the genus by DMW. Spathidexiaatripalpus sp. n., Spathidexiajuanvialesi sp. n., Spathidexiamarioburgosi sp. n., Spathidexialuisrobertogallegosi sp. n., Spathidexialuteola sp. n., Spathidexiahernanrodriguezi sp. n. and Spathidexiaaurantiaca sp. n. are all authored and described by Fleming and Wood. Minthodexiopsis Townsend is proposed by Wood as a new synonym of Spathidexia. A new combination proposed by Wood as a result of the new synonymy is S.flavicornis (Brauer & Bergenstamm) comb. n.

FIGURE 42. General morphology of Houghia sexmaculata sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia.

FIGURE 22. General morphology of Houghia fimbriata sp. nov. a. Dorsal habitus. b. Frontal view of head. c. Lateral habitus. d. Posterior view of male terminalia. e. Lateral view of male terminalia. f. Ventral view of sternite 5 + segments 7 + 8.