The continuity through the past 300 million years of key tropical sediment types, namely coals, evaporites, reefs and carbonates, is examined. Physical controls for their geographical distributions are related to the Hadley cell circulation, and its effects on rainfall and ocean circulation. Climate modelling studies are reviewed in this context, as are biogeographical studies of key fossil groups. Low-latitude peats and coals represent everwet climates related to the Intertropical Convergence Zone near the Equator, as well as coastal diurnal rainfall systems elsewhere in the tropics and subtropics. The incidence of tropical coals and rainforests through time is variable, being least common during the interval of Pangean monsoonal climates. Evaporites represent the descending limbs of the Hadley cells and are centred at 10° to 40° north and south in latitudes that today show an excess of evaporation over precipitation. These deposits coincide with the deserts as well as seasonally rainy climates, and their latitudinal ranges seem to have been relatively constant through time. Reefs also can be related to the Hadley circulation. They thrive within the regions of clear water associated with broad areas of downwelling which are displaced toward the western portions of tropical oceans. These dynamic features are ultimately driven by the subtropical high-pressure cells which are the surface signature of the subsiding branches of the Hadley circulation. Carbonates occupy the same areas, but extend into higher latitudes in regions where terrestrial surface gradients are low and clastic runoff from the land is minimal. We argue that the palaeo-latitudinal record of all these climate-sensitive sediment types is broadly similar to their environments and latitudes of formation today, implying that dynamic effects of atmospheric and oceanic circulation control their distribution, rather than temperature gradients that would expand or contract through time.

Carbon‐isotope analyses of fossil wood from the Middle Jurassic Ravenscar Group, Yorkshire, NE England, reveal a significant excursion toward light isotopic values (δ13C change of −3 to −4‰) at about the Aalenian‐Bajocian boundary (∼174 Ma). A positive carbon isotopic excursion is also shown for the middle Bajocian (∼170 Ma) but is less clearly defined. These isotopic patterns are very similar to the few published marine carbonate records available for this time, in particular one based on belemnites from the Hebrides basin, NW Scotland, and others from pelagic limestones in Italy. The similarity of the terrestrial and marine isotope curves is an indication that the observed isotopic signal is a global phenomenon. Through parts of the Ravenscar Group (the Scarborough Formation), supplementary data from bulk organic carbon and palynofacies analysis confirm that isotopic curves based on bulk analyses may be strongly influenced by the balance of terrestrial versus marine organic matter present in the samples. The negative isotope excursion at the Aalenian‐Bajocian boundary marks a change from charcoal to coal as the dominant preservational mode of the macroscopic wood fossils, which is interpreted here as a shift to a more continuously humid climate in the Early Bajocian. Upsection, charcoal once again becomes common, reflecting a return to more fire‐prone (presumably seasonally arid) environments in the middle Bajocian. Paradoxically, floral assemblages associated with the lithological unit in which the negative excursion occurs display characteristics that would normally be interpreted as adaptations to water stress brought about by relative aridity or salinity. Preliminary analyses of leaf stomatal densities show some evidence of raised pCO2 relative to background values at about the level of the negative excursion.

New Permian plant specimens are described from Prince Edward Island, Canada. They include attached specimens of leaf and stem genera Walchia and Tylodendron, enabling reconstruction of this Early Permian conifer. Although poorly preserved, the study of these floras extends our knowledge of diversity and climate conditions in the region. By placing these findings in a broader stratigraphic and geographic framework, we can document the phytogeographic and climate trends through the Carboniferous and Permian in the Maritimes Basin. Combined data on temporal trends in climate-sensitive sediments, as well as macrofloral and microfloral diversities, generally match the independently derived paleolatitudinal estimates. These show the region migrating from the southern subtropics across the Equator and into the northern subtropics between the Early Carboniferous and Early Permian. Evaporites and pedogenic carbonates, together with low-diversity floras, match its subtropical position in the Early Carboniferous. In contrast, coals are present in the Late Carboniferous, accompanied by high-diversity macro- and microfloral remains, when the region was on or near the Equator. However, the subsequent transition to pedogenic carbonates, eolian sands, and lower diversity floras is not matched by significant poleward latitudinal motion. We ascribe these changes to a decrease in moisture availability, as transgressions of epeiric seas became less frequent and finally stopped altogether, causing an increase of continentality in Euramerica.

Three new plant macrofossil assemblages were collected from Late Cretaceous and Paleocene fluvio-lacustrine sediments of the Pakawau and Kapuni groups in the northwest of the South Island, New Zealand. Palaeoenvironmental interpretations were made from each locality and palaeoclimate was deduced from the dicotyledonous angiosperm leaf component of each flora. A latest Cretaceous (Pakawau Bush Road locality) flora yielded 58 different dicotyledonous leaf forms; the two Paleocene collections, Ian’s Tip and Pillar Point Track, included 23 and 28 dicotyledonous leaf forms respectively. Quantitative palaeoclimate estimates were obtained using both Leaf Margin Analysis (LMA) and the Climate Leaf Analysis Multivariate Program (CLAMP). Temperature estimates suggest that there was a slight cooling from the latest Cretaceous into the early Paleocene in the northwest Nelson region of New Zealand, supporting similar Southern Hemisphere palaeoclimate findings from Antarctic data. Consistency in temperature estimates using different methods, including LMA, multivariate leaf morphological analysis (CLAMP), oxygen isotope data, regional versus local studies and global palaeoclimate models, suggests that the mean annual temperature for the Pakawau region in the latest Cretaceous was between 12 and 15°C. LMA produced temperature estimates between 6.5 and 8°C for the two Paleocene assemblages whereas CLAMP-produced estimates were slightly higher between 9 and 12.5°C

The Permian and Triassic represent a time of major global climate change from icehouse to hothouse conditions and significant (∼25°) northward motion of landmasses amalgamated in essentially one supercontinent, Pangea. The greatest of all mass extinctions occurred around the Permian-Triassic boundary (251 Ma), although there is no consensus regarding the cause(s). Recent studies have suggested a meteor impact and worldwide die-off of vegetation, on the basis of sparse local observations. However, new analyses of global Permian and Triassic plant data in a paleogeographic context show that the scale and timing of effects varied markedly between regions. The patterns are best explained by differences in geography, climate, and fossil preservation, not by catastrophic events. Caution should be exercised when extrapolating local observations to global-scale interpretations. At the other extreme, global compilations of biotic change through time can be misleading if the effects of geography, climate, and preservation bias are not considered.

The most recent global “icehouse‐hothouse” climate transition in earth history began during the Permian. Warmer polar conditions, relative to today, then persisted through the Mesozoic and into the Cenozoic. We focus here on two Permian stages, the Sakmarian (285–280 Ma) and the Wordian (267–264 Ma; also known as the Kazanian), integrating floral with lithological data to determine their climates globally. These stages postdate the Permo‐Carboniferous glaciation but retain a moderately steep equator‐to‐pole gradient, judging by the level of floral and faunal differentiation. Floral data provide a particularly useful means of interpreting terrestrial paleoclimates, often revealing information about climate gradations between “dry” and “wet” end‐member lithological indicators such as evaporites and coals. We applied multivariate statistical analyses to the Permian floral data to calibrate the nature of floral and geographical transitions as an aid to climate interpretation. We then classified Sakmarian and Wordian terrestrial environments in a series of regional biomes (“climate zones”) by integrating information on leaf morphologies and phytogeography with patterns of eolian sand, evaporite, and coal distributions. The data‐derived biomes are compared here with modeled biomes resulting from new Sakmarian and Wordian climate model simulations for a range of CO2 levels (one, four, and eight times the present levels), presented in our companion article. We provide a detailed grid cell comparison of the biome data and model results by geographic region, introducing a more rigorous approach to global paleoclimate studies. The simulations with four times the present CO2 levels (4×CO2) match the observations better than the simulations with 1×CO2, and, at least in some areas, the simulations with 8×CO2 match slightly better than those for 4×CO2. Overall, the 4×CO2 and 8×CO2 biome simulations match the data reasonably well in the equatorial and midlatitudes as well as the northern high latitudes. However, even these highest CO2 levels fail to produce the temperate climates in high southern latitudes indicated by the data. The lack of sufficient ocean heat transport into polar latitudes may be one of the factors responsible for this cold bias of the climate model. Another factor could be the treatment of land surface processes and the lack of an interactive vegetation module. We discuss strengths and limitations of the data and model approaches and indicate future research directions.

We use a climate model to simulate two intervals of Permian climate: the Sakmarian (ca. 280 Ma), at the end of the major Permo‐Carboniferous glaciation, and the Wordian (ca. 265 Ma). We explore the climate sensitivity to various levels of atmospheric CO2 concentration and to changes in geography and topography between the two periods. The model simulates large seasonality and high aridity in the continental interiors of both hemispheres for both periods. The northern summer monsoon weakens and the southern monsoon strengthens between the Sakmarian and the Wordian, owing to changes in geography and topography. The northern middle and high latitudes cool in winter, between the Sakmarian and Wordian, associated with northward shift of the continents. This high‐latitude cooling strengthens the winter westerlies and shifts the maximum storm‐track precipitation south. In the Southern Hemisphere, the winter westerlies weaken from the Sakmarian to the Wordian. Starting the simulations with no permanent ice fields (i.e., by assuming that the late Sakmarian postdates deglaciation) and imposing increased levels of atmospheric CO2 four times the present level, we find no tendency for reinitiation of major glaciation. Some permanent snow fields do develop in high southern latitudes, but these are primarily at high elevation. However, the combination of low CO2 levels (such as present‐day levels) and a cold summer orbital configuration produces expanded areas of permanent snow. The results are based on statistics derived from the final 5 yr of 20‐yr simulations. Paleoenvironmental indicators such as coal, evaporite, phosphate, and eolian sand deposits agree qualitatively with the simulated climate. The extreme cold simulated in high latitudes is inconsistent with estimates of high‐latitude conditions. Either the interpretation of observations is incorrect, the model is incorrect, or both; a possible model deficiency that leads to cold conditions in high latitudes is the relatively weak ocean‐heat transport simulated by the heat diffusion parameterization of the upper ocean model.

The most recent global icehouse‐hothouse climate transition in Earth history occurred in the Permian. Warmer polar conditions relative to today existed from the middle Permian through the Mesozoic and into the Cenozoic. We focus here on one particularly well-correlated middle Permian stage that postdated the deglaciation, the Wordian (267‐264 Ma), integrating floral and lithological data to determine Wordian climates globally. Paleobotanical data provide the best means of interpreting terrestrial paleoclimates, often revealing important information in the continuum between “dry” and “wet” end-member lithological indicators such as evaporites and coals. New statistical analyses of Wordian floras worldwide have enabled a greater understanding of original vegetation patterns and prevailing climate conditions. The derived climate interpretations are compared with new Wordian atmospheric general circulation model simulations. The model matches the data well in the tropics and northern high latitudes, but predicts colder conditions in southern high latitudes. We discuss possible reasons for this discrepancy.

An abstract is not available for this content so a preview has been provided. As you have access to this content, a full PDF is available via the ‘Save PDF’ action button.

An abstract is not available for this content so a preview has been provided. As you have access to this content, a full PDF is available via the ‘Save PDF’ action button.