The Anacacho Limestone was deposited during the Campanian and represents two depositional intervals, one of early Campanian and one of middle Campanian age. These two intervals correspond to periods of major eustatic sea level rise. This study focuses on the molluscan paleontology of the middle Campanian interval in the eastern part of the Anacacho exposure belt in Medina County, Texas. Molluscan assemblages in this area are indicative of inner to mid-shelf environments. No significant reef components are present. These eastern Anacacho deposits are interpreted to represent more offshore, deeper water environments than those to the southwest, where reef and lagoonal deposits have been reported.Analysis of the macrofossil components from these eastern localities has expanded the number of invertebrate species known from the Anacacho Limestone by nearly three-fold. This increase in diversity, based on a small amount of new work, suggests that many more taxa are yet to be identified, particularly in the western part of the exposure belt in Uvalde and Kinney Counties. This paper documents the bivalve and gastropod fauna, discussing and illustrating 24 bivalve taxa and 11 gastropod species. Two new bivalve species are named, Panopea anacachoensis new species and Spondylus siccus new species, and two potentially new gastropod species are identified but not named herein due to inadequate material. This paper expands the distribution of many eastern Gulf Coast and Atlantic Coast taxa westward into Texas and shows strong ties between the Anacacho fauna and that of the Campanian Tar Heel and Bladen Formations of the Black Creek Group in North Carolina. The taxonomic ties between these two areas probably reflect the thorough documentation of the North Carolina fauna, which is the best documented Campanian bivalve fauna in the Gulf or Atlantic Coast regions.
Upper Cretaceous limestone-shale couplets developed within the late transgressive stage of the Greenhorn cyclothem may be correlated from carbonate-dominated (basinal) sequences in central Kansas and Colorado westward to clastic cycles in southern Utah. Six such basinal couplets have been traced to corresponding upward-coarsening progradational cycles developed on the western margin of the Western Interior basin. In the central basin in Colorado and Kansas, these sedimentary cycles are represented by limestone-shale and maristone-shale couplets approximately 0.5-1.0 m in thickness. More calcareous parts of these couplets may be correlated westward into condensed, fossiliferous concretion and shell beds in proximal offshore lithofacies of Arizona and Utah. These concretion and shell beds are physically traceable farther landward (westward) into bioturbated, fossil-rich, transgressive lag deposits that bound 10- to 20-m-thick coarsening-upward progradational strand-plain deposits (parasequences) in southwestern Utah. Thus, the progradational phase of parasequence deposition correlates with accumulation of clay-rich sediment in the central basin, and the transgressive phase is characterized by reduced terrigenous input and deposition of carbonate-rich sediment.We consider Milankovitch-style orbital forcing of climate and tectonically induced fluctuations in rates of foredeep basin subsidence as possible forcing mechanisms for these basinwide events. Based on the widespread distribution of the limestone-shale couplets, as well as on estimated sedimentation rates and geochronology, it has been widely speculated that these carbonate cycles reflect Milankovitch cycles with periodicities on the order of 20 k.y. to 100 ky. If so, then stratigraphic data suggest that orbital forcing of climate affected eustasy and/or sediment input and biogenic production in the Western Interior basin. Alternatively, thrusting events in the Sevier orogenic belt may have produced episodic changes in the rates of foredeep basin subsidence and consequent changes in base level, which could have controlled the deposition of the Greenhorn parasequences and carbonate cycles. In either case, correlation of these units demonstrates a consistent basinwide sedimentary response to high-frequency base level or sediment input changes in the Western Interior epicontinental basin.
Upper Cretaceous rocks of the Kuskokwim Group are exposed in a large region of southwestern Alaska and are mainly composed of deformed turbidite deposits that contain few fossils other than inoceramid bivalves. This paper documents the taxonomy of the inoceramids in the Kuskokwim Group, develops an inoceramid biostratigraphy based on known ranges in other regions, and analyzes biogeographic patterns, paleoecology, and depositional history of the Kuskokwim Group.Most of the inoceramid bivalves present in the Kuskokwim Group are of Cenomanian and Turonian age, and an assemblage of species typical of late Turonian age rocks is particularly well developed. Only two localities appear to be as young as Santonian age. The following 16 species or subspecies are discussed and illustrated in detail:Birostrina tamuraiMatsumoto and Noda,Inoceramus virgatusSchlüter,I. pennatulusPergament,I. pictus minusMatsumoto,I.cf.I. yabeiNagao and Matsumoto,I.? sp. aff.I. costatusNagao and Matsumoto,I. hobetsensisNagao and Matsumoto,I. longealatusTröger,I. frechiFlegel,I. waltersdorfensis waltersdorfensisAndert,I.cf.I. waltersdorfensis hannovrensisHeinz,I. kuskokwimensisn. sp.,Mytiloidescf.M. opalensis(Böse),M. teraokai(Matsumoto and Noda),M.cf.M. incertus(Jimbo), andSphenoceramus naumanni(Yokoyama). In addition, a specimen with affinities toMytiloides striatoconcentricus carpathicus(Simionescu) and a specimen that may belong to theI. (Cremnoceramus?) rotundatus–I.(C.)erectuslineage are illustrated.Most of the taxa present in the Kuskokwim region are found in other regions of the North Pacific, particularly Japan and eastern Siberia, or are found throughout the Northern Hemisphere. Only one species,I. kuskokwimensisn. sp., is new and may be endemic. North Pacific taxa are predominant in the Kuskokwim region, but intervals near the Cenomanian–Turonian Stage boundary and in the upper Turonian contain taxa characteristic of Europe and the Western Interior basin of North America; some of these taxa have not been recorded previously in the North Pacific region. Turonian heteromorph ammonite assemblages associated with inoceramids in the finer grained facies of the Kuskokwim region are similar to those found in coeval rocks of Japan and Germany.The depositional area of the Kuskokwim Group can be broken into two northeast-trending subbasins, the Kuskokwim River subbasin to the northwest and the Mulchatna River subbasin to the southeast, connected by the Nushagak Hills corridor. Within the Kuskokwim River subbasin, deposition apparently started earlier in the north (middle Cenomanian) than in the south (late Cenomanian to early Turonian), and prograding deltaic sedimentation along the western margin also appears to have started earlier in the north. No marine fossils younger than latest Turonian to earliest Coniacian are known from the Kuskokwim River subbasin. The youngest fossils identified are Santonian in age and are from deep-water deposits in the Nushagak Hills corridor. Few fossils are known from the Mulchatna River subbasin and age control is limited.
High-resolution stratigraphic analysis of 18 sections spanning the Cenomanian–Turonian Stage boundary in the western interior of the United States has allowed determination of the magnitude and pattern of molluscan extinction and disruption. Composite range data from all sections show that the faunal turnover across the stage boundary occurs in a series of narrow stratigraphic zones, defined by multiple first and last occurrences, separated by intervals displaying little or no taxonomic turnover. Two of the apparent extinction steps (bottom and top of theNeocardioceras juddiiZone) may be intercontinentally developed. The additional steps apparently reflect cyclic changes in water mass and substrate characteristics in the western interior basin produced in response to orbital forcing of climate. An interval (ca. 10-100 k.y. duration) of changing community structure and general biotic deterioration is found below each of the two potentially intercontinentally developed extinction steps. The most affected mollusks were those having intercontinental distributions (ammonites and inoceramid bivalves), suggesting that disruption of planktotrophic larval dispersal may have played a role in increasing extinction and speciation rates near the C–T boundary. The nekto-benthic ammonites were affected earlier and to a greater degree than the pelagic forms, implying progressive upward expansion of the oxygen minimum zone preceding the stage boundary.