Evolution of population structure on islands is the result of physical processes linked to volcanism, orogenic events, changes in sea level, as well as habitat variation. We assessed patterns of genetic structure in the giant tortoise of the Aldabra atoll, where previous ecological studies suggested population subdivisions as a result of landscape discontinuity due to unsuitable habitat and island separation. Analysis of mitochondrial DNA (mtDNA) control region sequences and allelic variation at 8 microsatellite loci were conducted on tortoises sampled in 3 locations on the 2 major islands of Aldabra. We found no variation in mtDNA sequences. This pattern corroborated earlier work supporting the occurrence of a founding event during the last interglacial period and a further reduction in genetic variability during historical time. On the other hand, significant population structure recorded at nuclear loci suggested allopatric divergence possibly due to geographical barriers among islands and ecological partitions hindering tortoise movements within islands. This is the first attempt to study the population genetics of Aldabra tortoises, which are now at carrying capacity in an isolated terrestrial ecosystem where ecological factors appear to have a strong influence on population dynamics.

Predicting and redressing the threat of species extinction is not a success story. The science of estimating extinction rates and risk prediction is approached in a manner that is difficult to apply in the field, and yet current integrated ecosystem management programmes in many parts of the world, which are trying to place the conservation of species into sustainable community projects, need predictive tools for planning land use programmes. Such programmes involve US$100s of millions of multilateral and bilateral aid; many predicated on a site's biodiversity importance, the risks of extinction, sustainable extraction, production forms of land use, community livelihoods, water, and many others factors, but increasingly on ensuring that after the pump-priming funds are finished the programmes are both environmentally and economically sustainable (Swingland 2002, 2003, 2004; Swingland et al. 2003). [Swingland I.R. 2002. In: Swingland I.R., Bettelheim E.C., Grace J., Prance G.T. and Saunders L.S. (eds), Carbon Biodiversity, Conservation and Income: An Analysis of a Free Market Approach to Land-use Change and Forestry in Developing and Developed Countries. Philosophical Transactions Royal Society London A: Mathematical, Physical and Engineering Sciences, London; Swingland I.R. (ed.) 2003. Capturing Biodiversity and Conserving Biodiversity: The Market Approach. Earthscan, London; Swingland I.R. (ed.) 2004. CO2 e biodiversità. Un approccio integrato a favore del clima e del patrimonio naturale. Edizioni Ambiente, Milano, Italy, 296 pp.; Swingland I.R., Bettelheim E.C. and Niles J.O. 2003. In: Swingland I.R. (ed.), Capturing Biodiversity and Conserving Biodiversity: The Market Approach. Earthscan, London] This involves predictions of ‘what if?’ what if laws are changed to prevent over utilisation and prevent ‘The Tragedy of the Commons’ where land ownership is vested in the state and people degrade the environment, and instead institute private land ownership. In places like China and much of the Far East, biodiversity is over-exploited as nearly all their species are used for food, medicine and construction purposes, and private land ownership in rural areas is rudimentary or absent. Since most species extinction is anthropocentric, research on species extinction needs to be more accessible and focussed on global problems.

List of figures and Tables * About the Contributors * Preface Acknowledgements * List of Acronyms and Abbreviations * Introduction * Part 1: Carbon and Climate Change - Forests, Carbon and Global Climate * Changes in the Use and Management of Forests For Abating Carbon Emissions: Issues and Challenges Under the Kyoto Protocol * An Overview of a Free-Market Approach To Climate Change and Conservation * Potential Carbon Mitigation and Income in Developing Countries from Changes in Use and Management of Agricultural and Forest Lands * the Role of Multilateral Institutions * Electricity Generation: Options for Reduction in Carbon Emissions * Measuring, Monitoring and Verification of Carbon Benefits for Forest-Based Projects * Understanding and Managing Leakage in Forest-Based Greenhouse-Gas-Mitigation Projects * Part 2: Environmental Services - The Influence of Land-Use Change and Landscape Dynamics on the Climate System: Relevance to Climate-Change Policy Beyond the Radiative Effect of Greenhouse Gases * Economic, Biological and Policy Constraints on the Adoption of Carbon Farming in Temperate Regions * The Role of Sustainable Agriculture and Renewable-Resource Management in Reducing Greeenhouse-Gas Emissions and Increasing Sinks in China and India * Social Capital from Carbon Property: Creating Equity for Indigenous People * Species Survival and Carbon Retention in Commercially Exploited Tropical Rainforest * Animal Conservation, Carbon and Sustainability * Collateral Biodiversity Benefits Associated with 'Free-Market' Approaches to Sustainable Land Use and Forestry Activities * Developing Markets for Forest Environmental Services: an Opportunity for Promoting Equity While Securing Efficiency * Part 3: the Future Model - Carbon Sinks and Emissions Trading Under the Kyoto Protocol: a Legal Analysis * Protecting Terrestrial Ecosystems and the Climate Through a Global Carbon market * Designing a Carbon Market That Protects Forests in Developing Countries * Greenhouse-Gas-Trading Markets * Index

The global carbon cycle is significantly influenced by changes in the use and management of forests and agriculture. Humans have the potential through changes in land use and management to alter the magnitude of forest-carbon stocks and the direction of forest-carbon fluxes. However, controversy over the use of biological means to absorb or reduce emissions of CO(2) (often referred to as carbon 'sinks') has arisen in the context of the Kyoto Protocol. The controversy is based primarily on two arguments: sinks may allow developed nations to delay or avoid actions to reduce fossil fuel emissions, and the technical and operational difficulties are too threatening to the successful implementation of land use and forestry projects for providing carbon offsets. Here we discuss the importance of including carbon sinks in efforts to address global warming and the consequent additional social, environmental and economic benefits to host countries. Activities in tropical forest lands provide the lowest cost methods both of reducing emissions and reducing atmospheric concentrations of greenhouse gases. We conclude that the various objections raised as to the inclusion of carbon sinks to ameliorate climate change can be addressed by existing techniques and technology. Carbon sinks provide a practical available method of achieving meaningful reductions in atmospheric concentrations of carbon dioxide while at the same time contribute to national sustainable development goals.

Increased trade in non-timber forest products (NTFPs) has been promoted as one possible means to slow tropical deforestation by increasing the economic value of intact forest. A market survey of NTFPs occurring in the Capim River basin in eastern Amazonia, Brazil demonstrated that the reality for many smallholder communities in frontier and remote regions includes chronic transportation difficulties, high variability in fruit production, perishable products and lack of market expertise. In some communities, declining abundance of NTFPs due to logging and fire has resulted in a lack of forest products to even meet subsistence needs. In areas close to cities where transportation is assured and where forest clearing has eroded the natural occurrence of some valuable native NTFPs, smallholders who manage and successfully market native fruit and medicinal species are overcoming these obstacles. In frontier regions undergoing rapid transformation, however, decline in locally used and regionally marketed NTFPs currently pose detrimental consequences for communities. Findings suggest that an overemphasis on NTFP marketing has diverted attention from local livelihood, resource access and subsistence issues.

This paper describes the convergence of environmental and financial markets, reviews the evolution of market-based environmental programmes as an example of the seven-stage evolutionary process witnessed in a variety of markets and summarizes the emergence of greenhouse-gas-mitigation markets and their potential role in advancing land stewardship, biodiversity and other environmental services. Emissions trading has been developed to meet the demand to reduce pollution while avoiding economic disruption. Consistent with the seven-stage pattern of market evolution, the US programme to reduce the damage from acid rain established a standardized environmental commodity, developed 'evidence of ownership' necessary for financial instruments and provided the infrastructure to efficiently transfer title. The success of the system in reducing pollution at low cost has provided a model for other market-based environmental protection initiatives. The demand for cost-effective action to reduce the threat of climate change has initiated the same evolutionary process for markets to reduce greenhouse-gas emissions. Many of the land- and forest-management practices that can capture and store atmospheric CO(2) can also provide other environmental benefits, such as biodiversity preservation and enhanced water quality. The presence of a carbon-trading market will introduce a clear financial value for capture and mitigation of CO(2) emissions, thus introducing a new source of funding for land stewardship and forest rehabilitation. The market is now emerging through a variety of 'bottom-up' developments being undertaken through governmental, multilateral, private-sector and non-governmental-organization initiatives. The extension of markets to other emerging environmental issues is now underway, and the linkages between environmental sustainability and capital markets are being more deeply understood. The early evidence indicates that environmental sustainability can be compatible with maximization of shareholder value.

New incentives for protection and in situ use of forests and the services they provide raise hopes for the reversal of tropical and temperate deforestation. Past management of forests appropriated the rights of forest communities, providing incentives to convert natural forest into financial capital through logging, while destroying the underlying physical property. Carbon trading aims to provide a means to convert the forest property into financial capital, while protecting the physical property of forests, thereby providing new incentives for in situ forest management. The potential for carbon-emission trading as a contributor to these new incentives is tempered by concerns that it is another tool for capitalists to exploit the indigenous communities of the developing world. Estimates of annual emission trading amounting to US $200 billion raise alarm bells about the effect of such trade in the developing world. People are right to be concerned, as the history of exploitation of indigenous people, the appropriation of their rights, the loss of forests and their benefits is well documented. This exploitation resulted in the exclusion of forest communities from the basic tenets for development created by the wealth generated by traded property. However, one virtue of trade is that it can be made subject to constraints. Through international treaties and agreements, trade can be constrained and national governments obliged to observe the rules of trade. The value of tradable carbon credits will be discounted or invalid if they do not meet these criteria, providing all parties with strong incentives to achieve the necessary performance standards relating to both processes and contracts. For carbon trading to develop social capital from natural capital requires the admission of forest communities into the polity and management of forest resources. In this paper we argue for responsible carbon-emission trading based on the clear and appropriate definition of carbon entitlements, with the proviso that trading respects the rights and needs of indigenous people. We adopt this position as emissions trading now seems inevitable and there should be proper rules to control this trade where it affects forests and their inhabitants. It is imperative that the poor and indigenous people are not excluded from these systems. Trading systems and the property systems they depend on need to be more accountable, transparent and inclusive of those features which we propose.

Concern over the ever more rapid and widespread losses of biodiversity has instigated various remedial actions: whether in situ conservation, such as the establishment of protected areas, or ex situ, such as the conservation of germplasm in gene banks. In the past, such activities were funded and managed by the public sector; however, in recent years, public support has declined and this has spawned a growing interest in conservation opportunities that might arise from 'free-market' approaches to sustainable land use and management. The UN Convention on Biological Diversity (CBD) is the key framework for articulating policies and actions on biodiversity; however, progress in developing suitable economic and market incentives for biodiversity conservation and its sustainable use has been slow, with activities such as bioprospecting and ecotourism making some, albeit limited, headway. Given the United Nations Framework for Climate Change or United Nations Framework Convention on Climate Change's (UNFCCC's) high profile within the public and private sectors, there is some potential for using it to help advance CBD objectives and provide the much-needed economic incentives for conservation, through some of the market-based mechanisms presented under the Kyoto Protocol. Significant potential lies in the fact that many 'natural' forests and certain other ecosystems are both major stores of carbon and areas of valuable biodiversity. Thus, any attempt at conserving these areas has the potential to yield both carbon and biodiversity benefits. So far, however, the conservation of natural forests is not included in the Kyoto Protocol's definition of sinks. Instead the creation of sinks - through the establishment of fast-growing monocultures - may well lead to biodiversity losses, especially if partly degraded lands are cleared for this purpose. If real progress is to be made, our understanding of the relationship between land use and biodiversity benefits needs to be improved, and more appropriate proxies for biodiversity need to be developed. At the same time, we need to have a clear understanding of the precise nature of the potential synergies and be more able to identify possible jointaction opportunities that exist between the UNFCCC, the CBD and the Convention on Combating Desertification and other international trade and economic agreements.

An unequivocal, precise, and generally accepted definition of biodiversity does not exist. However the need for such a definition that is both scientifically sensible and universally applicable is imperative to help guide the design of policy and programs for the future, as well as to make critical decisions in the present. One of the many reasons for this state of affairs is that the definition of biodiversity affects objectives in national and regional research and conservation management, and in international funding priorities. Because so much is now formally invested in using the word biodiversity, its definition will continue to play a crucial role in both biodiversity conservation planning and public policy.

In the past decade much attention has focused on the role that genetics can play in the formation of management strategies in conservation. Here, we describe genetic diversity in the world's largest lizard, the Komodo dragon (Varanus komodoensis), examining the evolutionary relationships and population genetic history of the four islands in south-east Indonesia, which form the vast majority of its range. We identify distinct genetic groups for conservation. The population on the island of Komodo shows by far the largest values of genetic divergence and is proposed that it should be a separate conservation management unit. Other populations, surviving either on small islands with substantially reduced genetic variability, or in isolated patches, are identified as particularly vulnerable to stochastic threats and habitat loss. Our results provide an example of how data defining intraspecific levels of genetic divergence can provide information to help management plans, ensure the maintenance of genetic variability across populations and identify evolutionary potential within endangered species.

In recent years, representatives of more than 40 families of reptiles have been studied to understand how environmental parameters affect sex determination. In this review, we summarise the distribution and taxonomic pattern of sex determining mechanisms, outline the main hypotheses of the adaptive significance of temperature sex determination (TSD), and of skewed population sex ratios. We also examine the competing hypotheses of the physiological and molecular mechanisms involved in TSD.

The effect on vegetation communities of release from grazing by camels and goats has been investigated in the Baynunah region of Abu Dhabi emirate, in The United Arab Emirates, by the study of an exclosure established 11 years previously. Also the effect of sprinkle irrigation (in the absence of grazing) on the rangeland vegetation was investigated. Perennial species richness was significantly lower outside the exclosure compared with inside on both sand and gravel substrata. Annual species richness however was not significantly different. Perrenial percentage covers were lower outside the exclosure compared with inside, especially on sand substrata. The perennial grass Stipagrostis plumosa (L) showed the greatest difference in percentage cover in this respect. The species richness of annuals and perennials was not significantly different between irrigated and non-irrigated areas within the exclosure. The perennial percentage cover was much greater on sprinkle irrigated sand and gravel substrata. The perennial which benefited the most was Zygophyllum hamiense, Scweinf. Annual percentage cover was lower in irrigated quadrats. Suggestions are made about the effect of intensive grazing on the vegetation communities at Baynunah and on the effectiveness of irrigation as a tool for increasing the quality of the rangeland for livestock and wildlife.