Click to increase image sizeClick to decrease image size AcknowledgementsL.T. Ellis acknowledges the support of The Natural History Museum, London (BM). D. García-Avila thanks to Coordinación de la Investigación Científica (CIC) at Universidad Michoacana de San Nicolás de Hidalgo (UMSNH) and PROMEP (PTC-269) for the financial support for research on Bryophytes and for lab equipment. The work of I.V. Czernyadjeva was carried out within the framework of the institutional research project (no. 121021600184-6) of the Komarov Botanical Institute of the Russian Academy of Sciences. The study by E.A. Ignatova was supported by Lomonosov Moscow State University reserarch project no. 121032500090-7.L. Söderström, B. Cañiza and T. Pócs thank Lars Hedenäs for information about possible specimens in S, Katarina Stenman for making specimens in UME available, and Lil Stevens and Len Ellis for information on type material in BM.B. Espinoza-Prieto and J. J. Atwood thank Jennifer Kluse at the Shirley C. Tucker Herbarium (LSU) at Louisiana State University for providing the specimen loan. B. Espinoza-Prieto thanks the Center for Conservation and Sustainable Development of the Missouri Botanical Garden for funding through the Alwyn H. Gentry Fellowship and Steve Churchill (MO) for funding bryological exploration in central Peru through the Andean Bryophytes project; also thanks to Servicio Nacional Forestal (SERFOR, Ministry of Agriculture and Irrigation, Peru) for providing collection permits N°182-2018-MINAGRI-SERFOR-DGGSPFFS. W.R. Alvaro Alba acknowledges the Herbario Amazónico Colombiano (COAH), especially Nicolás Castaño and Wilson Rodriguez, for allowing access to the collection of non-vascular plants.The study by E. Yu. Kuzmina was carried out within the framework of the basic projects theme of the laboratory of lichenology and bryology of Komarov Botanical Institute RAS (“Flora of lichens and bryophytes of Russia and phytogeographically important regions”) № 121021600184-6. The study of E.A. Glazkova was carried out within the framework of the research project no. АААА-А 19-119031290052-1 (Vascular plants of Eurasia: systematics, flora, plant resources) of the Komarov Botanical Institute RAS. The field investigations of E.A. Glazkova were supported by the Complex Expedition “East Bastion — Kuril Ridge” of the Russian Geographical Society and the Expedition Center of the Russian Federation, and she thanks the organisers and all participants of the expedition. S. Ștefănuț acknowledges the support by project no. RO1567-IBB03/2022 through the Institute of Biology Bucharest of Romanian Academy. M. Burghardt acknowledges that his collection of Prionolejeunea clementinae was made under investigation permit 003-2018-IC-FLO-DPAP-MA of the Ecuadorian Ministry of Environment.V. Hugonnot thanks Heribert Köckinger for checking material of Tortella mediterranea. V. Sahu and A. K. Asthana are grateful to the Director, CSIR-National Botanical Research Institute for encouragement and providing the research facilities and to the forest department officials of Madhya Pradesh for kindly permitting the study area. Financial assistance from the CSIR under the project MLP 0042 is also thankfully acknowledged. These authors also thank the institutional ethics committee for granting institutional MS number CSIR-NBRI_MS/2022/11/02. J. J. Atwood acknowledges partial support from the U.S. National Science Foundation (Grant DEB 1655479).

Click to increase image sizeClick to decrease image size AcknowledgementsL.T. Ellis acknowledges the support of The Natural History Museum, London (BM) and is grateful to Jim Shevock (CAS) for loaning recent collections of Philippine Calymperaceae to BM for identification.The identification work of R. Gabriel and C. F. S. Picanço was partially supported by FEDER (85%) and by Azorean Public Funds (15%), through the Operational Program Azores 2020 under the project AZORESBIOPORTAL – PORBIOTA (ACORES-01-0145-FEDER-000072). M. Burghardt, J. V. Montoya and C. Villamarín acknowledge the investigation permits 003-2018-IC-FLO-DPAP-MA (MB 9895) and MAAE-DBI-CM-2021-0161 (MB 11253a) of the Ecuadorian Ministry of Environment under which the collections of Dicranella callosa were made. J. V. Montoya and C. Villamarín acknowledge financial support by Fondo para la protección de Agua (Quito) and Universidad de Las Américas (Project FGE.JMC.21.05), and M. Burghardt acknowledges financial support from the Universidad de las Americas (Project AGR.MB.19.03).W.R. Álvaro Alba and A.M Aponte Rojas are grateful to the Herbario Amazónico Colombiano (COAH), especially Dairon Cárdenas, and the Herbario Nacional Colombiano (COL), especially Julio Betancur and Jaime Uribe, for allowing access to the collections of non-vascular plants.The study of A. Mežaka was carried out within the framework of the postdoctoral research project “Epiphyte metapopulation dynamics in boreo-nemoral forest landscape” (no. 1.1.1.2/VIAA/3/19/469) in co-operation to Joint Stock company: “Latvia’s State Forests”. The study of Ansis Opmanis was supported by the Life project: “Threatened species in Latvia: improved knowledge, capacity, data and awareness” (LIFE19GIE/LV/000857). The work of O.M. Afonina was carried out within the framework of the institutional research project «Flora and taxonomy of algae, lichens and bryophytes in Russia and phytogeographically important regions of the world» # 121021600184-6. The work of D. Tubanova was conducted in the framework RSF 22-24-20132, https://rscf.ru/project/22-24-20132/.The contribution by F. Lara, A. L. Mateo Jiménez and I. Draper was supported by the Spanish State Research Agency (grant PID2020-115149GB-C21) and by Ministerio de la Juventud (Idelissa Bonelli de Calventi’s Dominicanan women scholarship 2021–22 to A. L. Mateo); Ministerio de Medio Ambiente y Recursos Naturales of The Dominican Republic is acknowledged for research permits.The contribution by P. Širka was financially supported by the Scientific Grant Agency of the Slovak Republic under Grant VEGA 1/0624/21.

In May 2021, the liverwort and hornwort flora of Colombia and Ecuador was finally published. After being delayed twice, certainly due to the Covid-19 pandemic, it was eagerly received and could be called an opus magnum of a long career in hepaticology. Having worked and published for over 50 years in this field, the author of this flora, S. Robbert Gradstein, is an accomplished bryologist on an international level and considered the foremost expert on Neotropical liverworts. This rather sizable book, with over 700 pages, is divided into several parts, starting with a brief introduction and a chapter on diversity and endemism. A classification precedes the taxonomic treatment, which, with about 620 pages, is the most extensive segment occupying the bulk of the book. A glossary, references, and an index to scientific names conclude the flora. The reviewer’s assessment of this book is from an Ecuadorian point of view; his knowledge of the Colombian aspects of this publication is limited. Evaluating such an extensive work like the present one is a tedious task, and hardly all observations can be referred to on a few pages; therefore, the comments are limited to some examples. In the abbreviations section, two of the twentyfour provinces of Ecuador are missing and do not appear throughout the book. In the case of Santa Elena, it might be understandable since probably neither collections nor reports of liverworts and hornworts from that province exist. For Santo Domingo de Los Tsáchilas, however, at least some liverwort species are documented; see Burghardt (2021) for a brief discussion. The classification follows Söderström et al. (2016) with minor deviations, like the distinction of Adelanthaceae and Jamesoniellaceae, the inclusion of Anastrophyllaceae in Scapaniaceae, and Southbyaceae in Arnelliaceae, among others. The taxonomic treatment comprises keys and descriptions of about 850 species, consisting of 830 liverworts and 20 hornworts, including the proposal of about a dozen new combinations, species, and subspecies. Furthermore, treatments of several genera like Acrobolbus, Lejeunea, and Lophocolea, among others, are revised. A broad morphological species concept is presented throughout the book, reflected by the proposal of about 100 new synonyms. Seventy plates illustrate almost all genera and about half of the species. Generally, the keys work well. However, the key to the Plagiochila species, including over 100 couplets, is very complex and, at times, challenging to use. Maybe this is owed to the sixty-five species involved and the extreme morphological plasticity of some common species, which key out several times. Nevertheless, it is unclear why the author did not break this key down into several shorter keys as he has done with Lejeunea, a similarly specious genus. There exists conflicting information on the status of Gongylanthus oniscoides (Spruce) Steph.

Acrobolbus mashpianus, a leafy liverwort of the family Acrobolbaceae, is described as new to science from the Chocó Biogeographical Region. It is characterized by strongly asymmetrical, deeply bilobed leaves with ciliate marginal teeth. The new species is illustrated and its relationships within the genus Acrobolbus are discussed. A key to the Ecuadorian species of Acrobolbus is provided.

While revising samples of Dialytrichia saxicola in LISU in preparation for IUCN Threatened Species of Europe assessments, (Sérgio & Porley 2019) we found a collection from Italy that matched the typical expression of this species.The population trend of this species was suspected to be increasing, favoured perhaps by eutrophication, and it was predicted that new areas of occurrence may be found (Sérgio & Porley 2019).It was therefore assessed as Least Concern, and considered native in Belgium, mainland France, Germany, Netherlands, mainland Portugal, Madeira, mainland Spain and Great Britain (Hodgetts & Lockhart 2020).Dialytrichia saxicola was first described from France as Dialytrichia mucronata (Brid.)Broth.var.fragilifolia Bizot & Cl.Roux, and subsequently raised to species rank as Dialytrichia fragilifolia (Bizot & Cl.Roux) F. Lara.It was later regarded as a variety of Dialytrichia mucronata, but taxonomic doubts remained pending further studies.More recently Barbula saxicola Lamy was shown to apply to this species and the transfer of Barbula saxicola to Dialytrichia necessitated the new combination D. saxicola (Lamy) M.J. Cano, since saxicola is the oldest available epithet

New national and regional bryophyte records, 70 L. T. Ellis, O. M. Afonina, M. H. B. Alia, M. Burghardt , B. Cabezudo, M. J. Cano, A. C. Cottet, J. Csiky, J. Deme, P. Erzberger, M. Evangelista, E. A. Glazkova, D. Gómez-González, J. Guerra, J. A. Jiménez, E. Yu. Kuzmina, N. S. Liksakova, M. I. Messuti, R. Natcheva, N. Norhazrina, J. P. Pantović, B. Papp, A. D. Potemkin , E. Rodríguez-Quiel, M. S. Sabovljević, D. Spitale, S. Ștefănuţ, N. Syazwana, M. G. Tossou and A. A. Vilnet t Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Komarov Botanical Institute of the Russian Academy of Sciences, St. Petersburg, Russia; Laboratoire de Botanique et Ecologie Végétale, Faculté des Sciences et Techniques, Université d’Abomey-Calavi, Cotonou, Benin; Grupo de Investigación en Biodiversidad, Medio Ambiente y Salud, BIOMAS / Facultad de Ingenierías y Ciencias Agropecuarias, Universidad de Las Américas, Quito, Ecuador; Área de Botánica, Facultad de Ciencias, Universidad de Málaga, Málaga, Spain; Facultad de Biología, Universidad de Murcia, Murcia, Spain; Instituto de Investigaciones en Biodiversidad y Medioambiente (INIBIOMA); Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET) – Universidad Nacional del Comahue (UNComahue), Quintral, Río Negro, Argentina; Department of Ecology, Institute of Biology, Faculty of Sciences, University of Pécs, Pécs, Hungary; Berlin, Germany; Universidade Estadual de Feira de Santana, Feira de Santana, BA – Brazil; Herbario UCH, Universidad Autónoma de Chiriquí, Panamá; Vicerrectoría de Investigación y Posgrado, Puebla, Mexico; Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, Sofia, Bulgaria; Department of Biological Sciences and Biotechnology, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia; Institute of Botany and Botanical Garden, Faculty of Biology, University of Belgrade, Belgrade, Serbia; Department of Botany, Hungarian Natural History Museum, Budapest, Hungary; Department of Botany, Institute of Biology and Ecology, Faculty of Science, Pavol Jozef Šafárik University in Košice, Košice, Slovakia; MuSe Museo delle Scienze, Corso del Lavoro e della Scienza, Trento, Italy; Institute of Biology Bucharest of Romanian Academy, Bucharest, Romania; PolarAlpine Botanical Garden-Institute of the Russian Academy of Sciences, Apatity, Murmansk Province, Russia

New national and regional bryophyte records, 67 L. T. Ellis, M. Alataş, W. R. Álvaro Alba, A. M. Charry Giraldo, V. Amatov, N. Batan, D. A. Becerra Infante, M. Burghardt, I. V. Czernyadjeva, E. Yu. Kuzmina, G. Ya. Doroshina, H. Erata, R. Garilleti , S. R. Gradstein, I. Jukonienė, S. Karaman Erkul, A. Keskin, T. Ezer, F. Lara , I. Draper , A. I. Maksimov, A. V. Mammadova, R. Natcheva, Cs. Németh, J. Pantović, M. S. Sabovljević, B. Papp, S. Poponessi, A. Cogoni, R. D. Porley, M. E. Reiner-Drehwald, A. Schäfer-Verwimp, A. Schmotzer, V. Šegota, A. Alegro, A. Rimac, S. Ştefănuț, E. Szurdoki, E. F. Vilk, V. M. Virchenko, R. J. Bijlsma and D. A. Callaghan Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Department of Plant and Animal Production, Munzur University, Vocational School of Tunceli, Tunceli, Turkey; Universidad Nacional de Colombia, Grupo de Investigación Biología de las Criptógamas de Colombia, Bogotá, Columbia; Faculty of Science and Art, Biology Department, Recep Tayyip Erdoğan University, Rize, Turkey; Macka Vocational School, Karadeniz Technical University, Trabzon, Turkey; Universidad Pedagógica y Tecnológica de Colombia, Grupo de Investigación Biología para la Conservación, Columbia; Carrera de Ingeniería Agroindustrial y Alimentos, Facultad de Ingeniería y Ciencias Agropecuarias, Universidad de Las Américas, Quito, Ecuador; Komarov Botanical Institute of the Russian Academy of Sciences, St. Petersburg, Russia; Bayramiç Vocational School, Çanakkale Onsekiz Mart University, Çanakkale, Turkey; Departamento de Botánica y Geología, Facultad de Farmacia, Universidad de Valencia, Burjassot, Spain; Department of Systematics, Biodiversity and Evolution of Plants, Albrecht von Haller Institute, University of Göttingen, Göttingen, Germany; Nature Research Centre, Vilnius, Lithuania; Aksaray University, Faculty of Science, Department of Biology, Aksaray-Turkey; Faculty of Science, Department of Biology, Niğde Ömer Halisdemir University, Niğde, Turkey; Departamento de Biología (Botánica), Facultad de Ciencias, Universidad Autónoma de Madrid, Madrid, Spain; Centro de Investigación en Biodiversidad y Cambio Global, Madrid, Spain; Institute of Biology, Karelian Research Center of the Russian Academy of Sciences, Petrozavodsk, Russia; Institute of Botany, Azerbaijan National Academy of Sciences, Baku, Azerbaijan; Institute of Biodiversity and Ecosystem Research Bulgarian Academy of Sciences, Sofia, Bulgaria; Institute of Ecology and Botany, Vácrátót, Hungary; Institute of Botany and Botanical Garden, Faculty of Biology, University of Belgrade, Belgrade, Serbia; Botanical Department, Hungarian Natural History Museum, Budapest, Hungary; Department of Environmental and Life Science, University of Cagliari, Sardinia, Italy; Cera dos Pomares, Aljezur, Portugal; Wacholderweg 24, 37079, Göttingen, Germany; Mittlere Letten 11, Herdwangen-Schönach D-88634, Germany; Bükk National Park Directorate, Hungary; Herbarium Croaticum, Department of Botany, University of Zagreb, Zagreb, Croatia; Institute of Biology Bucharest of Romanian Academy, Bucharest, Romania; Faculty of Primary and Pre-School Education, Department of Natural Sciences, Eötvös Lóránd University, Budapest, Hungary; Institute of Biological Problems of the North of the Far Eastern Branch of RAS, Magadan, Russia; Department of Phycology, Lichenology and Bryology, Institute of Botany, National Academy of Sciences of Ukraine, Kyiv, Ukraine; Wageningen Environmental Research, Wageningen University & Research, Wageningen, The Netherlands; Bryophyte Surveys Ltd, Bristol, UK

Lepicolea ochroleuca is newly recorded for Ecuador and Plagiochila cuneata is new to continental Ecuador, previously known only from the Galapagos Islands. Seventy-seven new provincial records of 62 liverwort species distributed among 15 provinces are presented. The uneven knowledge of the distribution of liverworts in Ecuador is briefly discussed and some observations on collections by non-bryologists are made.

A new species of Symphyogyna, S. caduciloba, is described, based on the combination of ascending growth form, deeply lobate thallus, variable lobe tips with a small, 1–2-celled apical tooth which is sometimes replaced by a slime papilla, and caducous thallus lobes. Relations with similar species are discussed and a key to the lobate species of Symphyogyna in Ecuador is presented.

1. Aloina brevirostris (Hook & Grev.) Kindb. Contributor. M. Puglisi Sicily. Rocca di Entella, plateau (municipality of Contessa Entellina), 37°46'24.1N, 13°06'58.4E, 483 m a.s.l., 4 March 2019, ...

1. Anthoceros tristanianus J.C.Villarreal, J.J.Engel & J.Vaňa Contributors. B. Espinoza-Prieto and J. Opisso Peru. Antabamba Province, Dpto. Apurimac: Juan Espinoza Medrano district, Paraje Ccancca...

A preliminary list of the bryophytes of the Mashpi Ecological Reserve is presented and 112 species and one variety of liverworts, 41 species of mosses and one species of hornworts are reported. Nine liverworts (Drepanolejeunea spinosa, Harpalejeunea subacuta, Neurolejeunea sastreana, Plagiochila eggersii, P. rudischusteri, Prionolejeunea magnistipula, P. trachyodes, Trichocolea filicaulis and Zoopsidella integrifolia) and one moss (Cyclodictyon subtortifolium) are newly recorded for Ecuador. Furthermore, 70 liverworts, 19 mosses and one hornwort are registered for the first time for the Province of Pichincha.

Leptoscyphus Mitten (1851: 358), a medium-sized genus of the Lophocoleaceae, is mainly distributed in tropical and temperate South America, Africa and Australasia, and is generally characterized by the brownish gametophyte colour and a laterally compressed perianth with a more or less bilabiate mouth due to the reduction of the ventral lobe (Vanderpoorten et al. 2010). 15 species are reported to occur in Ecuador (Gradstein & Benitez 2017). During revision of recent collections in the National Herbarium of Ecuador (QCNE) an undescribed species came to our attention which we here describe and illustrate.

Two liverworts (Gymnocoleopsis cylindriformis and Schiffneriolejeunea polycarpa) are reported as new to Ecuador as well as 69 new provincial records. Furthermore, corrections are presented for some previously published records.

1. Aulacopilum glaucum WilsonContributors. P. Srivastava, V. Sahu and A. K. AsthanaIndia. Manipur, Imphal, Bishnupur, Leimaram, near Sadu Chiru Falls, 24°44′21.3″N, 093°44′56.3″E, epiphytic, 1111 m...

L. T. Ellis, L. A. Amélio, D. F. Peralta, M. Bačkor, E. Z. Baisheva, H. Bednarek-Ochyra, M. Burghardt, I. V. Czernyadjeva, S. S. Kholod, A. D. Potemkin , A. Erdağ, M. Kırmacı, V. E. Fedosov, M. S. Ignatov, D. E. Koltysheva, J. R. Flores, E. Fuertes, M. Goga, S.-L. Guo, W. K. Hofbauer, M. Kurzthaler, H. Kürschner, O. I. Kuznetsova, M. Lebouvier, D. G. Long, Yu. S. Mamontov, K. M. Manjula, C. N. Manju, B. Mufeed, F. Müller, M. C. Nair, M. Nobis, N. Norhazrina, M. Aisyah, G. E. Lee, M. Philippe, D. A. Philippov, V. Plášek, Z. Komínková, R. D. Porley, Yu. A. Rebriev, M. S. Sabovljević, A. M. de Souza, E. B. Valente, D. Spitale, P. Srivastava, V. Sahu, A. K. Asthana, S. Ştefănuţ, G. M. Suárez, A. A. Vilnet, K.-Y. Yao and J.-Ch. Zhao The Natural History Museum, Cromwell Road, London, SW7 5BD, UK; Instituto de Botânica, São Paulo, Brazil; Department of Botany, Institute of Biology and Ecology, Pavol Jozef Šafárik University, Košice, Slovakia; Ufa Institute of Biology, Subdivision of Ufa Federal Research Centre of Russian Academy of Sciences, Ufa, Russia; W. Szafer Institute of Botany, Polish Academy of Sciences, Kraków, Poland; Carrera de Ingeniería Agroindustrial y Alimentos, Facultad de Ingeniería y Ciencias Agropecuarias, Universidad de Las Américas, Pichincha, Ecuador; Komarov Botanical Institute of the Russian Academy of Sciences, St. Petersburg, Russia; Aydın Adnan Menderes Üniversitesi, Kepez-Aydin, Turkey; Lomonosov Moscow State University, Russia; Finnish Museum of Natural History, University of Helsinki, Helsinki, Finland; Departamento de Biodiversidad, Ecología y Evolución, Universidad Complutense, Madrid, España; Institute of Biology and Ecology, Pavol Jozef Šafárik University, Košice, Slovakia; Core Facility Cell Imaging and Ultrastructure Research, University of Vienna, Vienna, Austria; East China Normal University, Shanghai, People’s Republic of China; Fraunhofer-Institut für Bauphysik IBP, Holzkirchen, Germany; Freie Universität Berlin, Institut für Biologie, Systematische Botanik und Pflanzengeographie, Berlin, Germany; Tsitsin Main Botanical Garden, Russian Academy of Sciences, Moscow, Russia; CNRS UMR 6553, Université de Rennes 1, France; Royal Botanic Garden Edinburgh, Edinburgh, UK; Botanical Garden-Institute, Far Eastern Branch of Russian Academy of Sciences, Vladivostok, Russia; PG & Research Department of Botany, The Zamorin’s Guruvayurappan College, Kozhikode, Kerala, India; Institut für Botanik, Technische Universität Dresden, Dresden, Deutschland; PG & Research Department of Botany, Govt. Victoria College, Palakkad, Kerala, India; Jagiellonian University, Kraków, Poland; Faculty of Science and Technology, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia; Université Lyon-1 et UMR CNRS 5023 ‘Ecologie des Hydrosystèmes Naturels et Anthropisés’, Villeurbanne, France; Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences, Borok, Yaroslavl Region, Russia; University of Ostrava, Ostrava, Czech Republic; Cera dos Pomares, Aljezur, Portugal; Southern Scientific Center of the Russian Academy of Sciences, Rostov-on-Don, Russia; Institute of Botany and Botanical Garden, Faculty of Biology, University of Belgrade, Belgrade, Serbia; Departamento de Biologia, Universidade Estadual de Feira de Santana, Feira de Santana, BA—Brazil; MuSe Museo delle Scienze, Corso del Lavoro e della Scienza, Trento, Italy; Bryology Laboratory, CSIR–National Botanical Research Institute, Lucknow, India; Institute of Biology Bucharest of Romanian Academy, Bucharest, Romania; Unidad Ejecutora Lillo (UEL; FML-CONICET), Miguel Lillo 251, (4000) S.M. de Tucumán, Argentina; Facultad de Ciencias Naturales e Instituto Miguel Lillo, Universidad Nacional de Tucumán, Miguel Lillo 205, (4000) S.M. de Tucumán, Argentina; PolarAlpine Botanical Garden, Kola Science Centre, Russian Academy of Sciences, Murmansk Province, Russia; Endemic Species Research Institute, Nantou, Taiwan; Hebei Normal University, Shijiazhuang, People’s Republic of China

1. Amphidium lapponicum (Hedw.) Schimp.Contributor. R. OchyraPoland. Western Carpathians, Western Beskidy Mountains: Beskid Żywiecki Range, Babia Gora massif, on the northern slope of Diablak peak,...

Ecuador has a very diverse bryophyte flora with about 950 species of mosses and 700 of liverworts and hornworts. Nevertheless, the distribution of the species within the country remains incompletely explored and many species are only known from very few collections. This paper presents new additions to the liverwort and hornwort flora of Ecuador. The hornwort Notothylas vitalii and the liverworts Blepharostoma trichophyllum, Frullania setigera, Isopaches bicrenatus, Platycaulis renifolia and Symphyogyna apiculispina are new to the country, four species are new to southern Ecuador, eighteen are new to the province of Loja, ten are new to Napo, six are new to Pichincha and one species is new to the provinces of El Oro and Carchi respectively. The record of Notothylas vitalii is the first one outside Brazil and constitutes the first record of the genus Notothylas from mainland Ecuador. The rare monospecific genus Platycaulis was previously known only from the type locality in the Venezuelan Andes, and the holarctic Isotaches bicrenatus in the tropics only from two mountains in southeastern Brazil. Several new records were gathered in very humid and bryophyte-rich, wind-stricken, foggy superpáramo vegetation in the Páramo de la Virgen (Napo) at 4100-4300 m. For all species notes on their geographical distribution and habitats in Ecuador as well as their world range are provided.

The liverwort Odontoschisma engelii Gradst. & Burghardt sp. nov. (Cephaloziaceae) is described from páramo vegetation of southern Ecuador and Venezuela. The new species is a member of Odontoschisma section Denudatae R. M. Schust. and differs from all known species of the genus by the strongly elongated leaves.