The North Sea Basin has been covered by ice sheets originating from both the British Isles and Scandinavia at multiple times during the Pleistocene. The Witch Ground Basin (WGB) in the central northern North Sea is a critical location in terms of interpreting Late Pleistocene glacial to glacimarine history of the North Sea since it was the location of the Witch Ground Ice Stream that was active on multiple occasions during the Mid to Late Pleistocene. We map five mega-scale glacial lineation flowsets corresponding to the changing ice flow direction of the Witch Ground Ice Stream and investigate the sedimentological fingerprint and corresponding subglacial depositional processes of this palaeo-ice stream. We show that sorted sand layers within a subglacial traction till represent periodic hydraulic jacking and ice-bed decoupling at the base of the Witch Ground Ice Stream. In contrast to previous studies that have described glacitectonites deposited below the most recent grounded ice in the WGB, we present analysis of sediment cores that recovered primarily massive diamictons without any obvious deformation structures. The most recent ice cover in the WGB (similar to 18-16 ka) was thought to have been sourced from a localized ice cap over Orkney and Shetland. The presence of chalk clasts sourced from NW of the WGB described in this study from the stratigraphically youngest till confirms this interpretation. The transition from subglacial to glacimarine deposition, while acoustically well defined (from opaque to laminated acoustic units), appears surprisingly uniform in the recovered sediment cores, but can be differentiated based on a change in colour including mottling and banding, presence of whole intact shells, and the increased number of silt and sand lenses. C-14 dating of glacimarine muds indicate high sedimentation rates of between 80 and 260 cm ka(-1). The transition from glacimarine to marine deposition is represented by a comparative decrease in sedimentation rate and deposition of Holocene age sandy mud. This study demonstrates a highly dynamic Witch Ground Ice Stream in the northern North Sea during the Late Pleistocene with evolving subglacial hydrology and depositional processes at the ice stream bed that left a distinct geomorphological and sedimentological fingerprint within the WGB.
Floristic homogenization - an increase in plant similarity within a given region - threatens biodiversity. By studying the taxonomic similarity of the floras of South Pacific islands over the past 5,000 years, we find that initial human settlement was probably a major driver of floristic homogenization.
The increasing similarity of plant species composition among distinct areas is leading to the homogenization of ecosystems globally. Human actions such as ecosystem modification, the introduction of non-native plant species and the extinction or extirpation of endemic and native plant species are considered the main drivers of this trend. However, little is known about when floristic homogenization began or about pre-human patterns of floristic similarity. Here we investigate vegetation trends during the past 5,000 years across the tropical, sub-tropical and warm temperate South Pacific using fossil pollen records from 15 sites on 13 islands within the biogeographical realm of Oceania. The site comparisons show that floristic homogenization has increased over the past 5,000 years. Pairwise Bray–Curtis similarity results also show that when two islands were settled by people in a given time interval, their floristic similarity is greater than when one or neither of the islands were settled. Importantly, higher elevation sites, which are less likely to have experienced human impacts, tended to show less floristic homogenization. While biotic homogenization is often referred to as a contemporary issue, we have identified a much earlier trend, likely driven by human colonization of the islands and subsequent impacts.
Introduction Islands of the Southwest Pacific are exposed to geologic and climate-related disturbances that occur on a range of timescales and which probably affect, to varying degrees, their terrestrial ecosystems. Over the past ∼1100 years we know of two major events in the region: the Kuwae eruption which is thought to have occurred ∼500 cal. years BP and a shift to drier conditions which began ∼1100 cal. years BP. Methods We investigated terrestrial and lacustrine ecosystem responses to these events and also to a changing fire regime, likely human-caused, using a multi-proxy (C/N, charcoal, chironomids, pollen, and tephra) record from Lake Emaotul, Efate, Vanuatu. Results Tephra from the Kuwae eruption was found across a 6 cm layer which our age-depth model suggests was deposited 650–510 cal. years BP (95% confidence). Forest and chironomid community turnover increased during the wet-dry shift 1100–1000 cal. years BP; subsequently, chironomid turnover rates decreased again within <135 years and vegetation had partially (but not fully) recovered after ∼80 years. Following Kuwae volcanic tephra deposition, vegetation turnover increased again, reflecting a reduction in small trees and shrubs and an increase in grasses. Subsequently, the forest vegetation did not regain its previous composition, whereas chironomid community composition remained fairly stable before and after tephra deposition. Within the last ∼90 years, enhanced local burning drove another increase in vegetation turnover. Discussion Terrestrial and freshwater ecosystems in Efate are sensitive to changes in hydroclimate, volcanism, and anthropogenic fires, although to different degrees; while recent human impacts are often obvious, volcanic eruptions and climatic shifts have also structured Pacific-island ecosystems and will continue to do so.
Cabo Verde remained uninhabited until 1460 CE, when European sailors founded a settlement in Santiago, and soon after in Fogo island. The degree to which different island ecosystems in Cabo Verde have been transformed by humans remains uncertain because of a scarcity of historical information and archaeological evidence. Disentangling these processes from natural ones is complicated in islands with a history of volcanic impacts and other natural hazards. In this paper, we apply microfossil (pollen, non-pollen palynomorphs and phytoliths) and sedimentological analyses (granulometry, X-ray diffraction, loss on ignition and tephrostratigraphy) to a 2-m sediment sequence deposited in a scoria cone from 4100 cal year BP (calibrated years before 1950 CE) to the present. The organic-rich basal sediments indicate that between 4100 and 2600 cal year BP the pre-settlement landscape of Fogo was an open grassland, where fire was infrequent and/or small-scale. An increase in volcanic glass deposition after 2600 cal year BP, peaking ca. 1200 cal year BP, suggests that there was a progressive activation of Fogo’s volcanic activity, contemporaneous with increased fire frequency and erosion pulses, but with little impact on local grassland vegetation. While dating uncertainty is high, the first evidence of intensive local land use by early settlers was in the form of cultivation of Zea mays, abundant spores of coprophilous fungi (i.e. Sporormiella), and peaks in charcoal concentrations between 800 and 400 cal year BP. This was followed by large increases in pollen from pigeon pea ( Cajanus), a diverse array of exotic trees ( Cupressus, Grevillea), and invasive shrubs ( Lantana). The introduction of these taxa is part of recent human effort to ‘tame’ this steep, dry and hazardous island by reducing erosion and providing firewood. An important outcome of these efforts, however, is a loss of fragile native biodiversity.
Human-mediated changes in island vegetation are, among others, largely caused by the introduction and establishment of non-native species. However, data on past changes in non-native plant species abundance that predate historical documentation and censuses are scarce. Islands are among the few places where we can track human arrival in natural systems allowing us to reveal changes in vegetation dynamics with the arrival of non-native species. We matched fossil pollen data with botanical status information (native, non-native), and quantified the timing, trajectories and magnitude of non-native plant vegetational change on 29 islands over the past 5000 years. We recorded a proportional increase in pollen of non-native plant taxa within the last 1000 years. Individual island trajectories are context-dependent and linked to island settlement histories. Our data show that non-native plant introductions have a longer and more dynamic history than is generally recognized, with critical implications for biodiversity baselines and invasion biology.
Sa similar to o Tome ' (Gulf of Guinea, Central Africa) is a 854 km2 tropical island that had a pivotal role in early European colonial expansion through the Atlantic between the 15th and 16th centuries. Historical sources suggest that native vegetation has been heavily impacted since human arrival (1470 CE) due to monoculture economies and the introduction of mammals and plants, some of which now have established wild populations. The Afro-montane forest of Sa similar to o Tome ', located above 800 m.a.sl., is particularly rich in endemic plant species and has remained relatively unaffected by direct human impacts. Here, we explore how environmental change influenced this forest through the study of a sedimentary sequence from the volcanic crater of Lagoa Ame ' lia (1340 m a.s.l.), a palustrine system located at the boundary between submontane (800-1400 m a.s.l.) and mist forest (above 1400 m a.s.l.). We used fossil pollen, non-pollen palynomorphs, sedimentology and charcoal to determine forest dynamics from the Late Pleistocene to the present. From 14,000 to 12,500 cal yr BP the forest was dominated by taxa from higher altitudes, adapted to cooler and drier climates (e.g. Afrocarpus mannii trees and Psychotria nubicola). After 12,500 cal yr BP, a potential uphill migration was identified by an increase in taxa like the trees Symphonia globulifera and Craterispermum cerinanthum. From 11,200 cal yr BP through the rest of the Holocene taxa from lower altitudes became dominant (e.g. Prunus africana, Polyscias, and Sabicea), except at c. 8500 cal yr BP when rapid cooling led to forest opening. Charcoal showed that fires were frequent during the Late Pleis-tocene (14,000 to 11,200 cal yr BP), becoming rare during the Holocene until anthropogenic fires started at c. 220 cal yr BP. Other recent anthropogenic impacts detected in Lagoa Ame ' lia included the appearance of pollen of introduced plant species (e.g., Cestrum), and the increase in pollen of economically important species (Elaeis guineensis, Zea mays) and in fungal spores related to introduced herbivores. Our results reveal that climate changed the altitudinal distribution of the Afromontane forest in Sa similar to o Tome ' during the Late Pleistocene, as observed on the African continent, and that this ecosystem was also strongly impacted by human arrival, through fire, farming, and introduced species.
Here, we investigate Mid- to Late-Holocene vegetation changes in low-lying coastal areas in Tonga and how changing sea levels and recurrent volcanic eruptions have influenced vegetation dynamics on four islands of the Tongan archipelago (South Pacific). To investigate past vegetation and environmental change at Ngofe Marsh ('Uta Vava'u), we examined palynomorphs (pollen and spores), charcoal (fire), and sediment characteristics (volcanic activity) from a 6.7-m-long sediment core. Radiocarbon dating indicated the sediments were deposited over the last 7700 years. We integrated the Ngofe Marsh data with similar previously published data from Avai'o'vuna Swamp on Pangaimotu Island, Lotofoa Swamp on Foa Island, and Finemui Swamp on Ha'afeva Island. Plant taxa were categorized as littoral, mangrove, rainforest, successional/ disturbance, and wetland groups, and linear models were used to examine relationships between vegetation, relative sea level change, and volcanic eruptions (tephra). We found that relative sea level change has impacted vegetation on three of the four islands investigated. Volcanic eruptions were not identified as a driver of vegetation change. Rainforest decline does not appear to be driven by sea level changes or volcanic eruptions. From all sites analyzed, vegetation at Finemui Swamp was most sensitive to changes in relative sea level. While vegetation on low-lying Pacific islands is sensitive to changing sea levels, island characteristics, such as area and elevation, are also likely to be important factors that mediate specific island responses to drivers of change.
This study provides a high-resolution reconstruction of the vegetation of the Argive Plain (Peloponnese, Greece) covering 5000 years from the Early Bronze Age onwards. The well dated pollen record from ancient Lake Lerna has been interpreted in the light of archaeological and historical sources, climatic data from the same core and other regional proxies. Our results demonstrate a significant degree of human impact on the environments of the Argive Plain throughout the study period. During the Early Bronze Age evidence of a thermophilous vegetation is seen in the pollen record, representing the mixed deciduous oak woodland of the Peloponnesian uplands. The plain was mainly used for the cultivation of cereals, whereas local fen conditions prevailed at the coring site. Towards the end of this period an increasing water table is recorded and the fen turns into a lake, despite more arid conditions. In the Late Bronze Age, the presence of important palatial centres modified the landscape resulting in decrease of mixed deciduous oak woodland and increase in open land, partly used for grazing. Possibly, the human management produced a permanent hydrological change at Lake Lerna. From the Archaic period onwards the increasing human pressure in association with local drier conditions caused landscape instability, as attested by a dramatic alluvial event recorded in the Pinus curve at the end of the Hellenistic Age. Wet conditions coincided with Roman times and favoured a forest regeneration pattern in the area, at the same time as we see the most intensive olive cultivation in the pollen record. The establishment of an economic landscape primarily based on pastures is recorded in the Byzantine period and continues until modern times. Overgrazing and fires in combination with arid conditions likely caused degradation of the vegetation into garrigue, as seen in the area of the Argive Plain today.
Using diatoms, pollen, and geochemistry, we explore human habitation around Lina myr, Gotland, in relation to shore displacement. Archeological evidence has shown that Lina myr was an important area for its prehistoric human inhabitants. We investigate if and when Lina myr was connected to the sea and could therefore have been part of an inland water system useful for transport. A chronology was based on 14 C AMS dating of terrestrial macrofossils and bulk sediments with dates ranging between 9100 and 2360 cal. yr BP. The initiation of the Littorina transgression was dated to 8500 cal. yr BP. A twofold pattern for the maximum sub-phase of the Littorina Sea is suggested from 8100 to 7500 cal. yr BP and from 6500 to 6000 cal. yr BP. The onset of cultivation and grazing was indicated by the presence of Hordeum and Plantago lanceolata in the pollen record during the Late Neolithic, at about 4580 cal. yr BP. During this time sea level was relatively higher than today and the Lina myr basin was connected with the Littorina Sea, which it continued to be until isostatic uplift caused it to become isolated at about 3820 cal. yr BP. After about 3000 cal. yr BP, human-made landscape changes intensified, grasslands increased, and shrublands decreased.
Understanding past hydroclimate variability and related drivers is essential to improve climate forecasting capabilities especially in areas with high climatic sensitivity, such as the Mediterranean. This can be achieved by using a broad spectrum of high resolution, multiple proxy records which can also allow us to assess linkages between regional hydroclimate variability and shifts in the large-scale atmospheric patterns. Here, we present a multiproxy reconstruction of the central-eastern Mediterranean hydro climate changes over the last 3600 years based on a sediment core from the Gialova Lagoon, a shallow coastal ecosystem in SW Peloponnese, Greece. Our combined dataset consists of the distribution and compound-specific carbon and hydrogen isotope (delta C-13 and 8D) composition of n-alkanes, bulk organic matter properties and X-ray fluorescence (XRF) core scanning data. This approach was complemented with a semi-quantitative analysis of plant remains in the core. The results indicate a high contribution of local aquatic vegetation to organic matter. Large delta C-13 variations in predominantly aquatic plant-derived mid-chain alkanes (C23-23) mainly reflect changes in the aquatic plant abundance and their carbon source. Our data suggest that higher delta C-13(23-25) values (up to 19 parts per thousand) largely correspond to expansion of aquatic vegetation during wet and/or cold periods causing carbon-limiting conditions in the water and assimilation of isotopically-enriched bicarbonate by the plants. The 8D records of the individual n-alkanes (C-17 to C-31) exhibit a nearly identical pattern to each other, which implies that they all reflect changes in the source water isotope composition, driven by hydroclimate variability. In addition, the 8D profiles are consistent with the XRF data with both proxies being driven by a common hydroclimate signal. We observe two major shifts from dry and/or warm periods at ca 3600-3000 cal BP and ca 17001300 cal BP to wet and/or cold episodes at ca 3000-2700 cal BP and ca 1300-900 cal BP. The period ca 700-200 cal BP is the wettest and/or coldest in our record and coeval with the Little Ice Age. The climatic fluctuation reported in this study can be explained by the relative dominance of high-latitude (e.g. North Atlantic Oscillation during winters) and the low-latitude atmospheric patterns (Intertropical convergence zone, Subtropical High and the effects of Asian monsoons during summers) which suggests an Atlantic-Mediterranean-Monsoon climate link in this area for the late Holocene. (C) 2018 Elsevier Ltd. All rights reserved.