Drought stress (DS) is a major environmental factor impacting plant growth and productivity on a global scale. Due to rapidly changing climatic conditions, DS is proving to be the most detrimental among various stresses. It poses a significant threat to agricultural sustainability by hindering the growth and yield of agriculturally important crops worldwide. Plants have developed various adaptive mechanisms, including physio-biochemical changes, secondary metabolism, and genetic regulation, to cope with such challenging conditions. This review provides a comprehensive analysis of how DS influences physiological processes such as photosynthesis, and stomatal conductance, alongside biochemical adjustments. Furthermore, this review investigates the alterations in secondary metabolism, emphasizing the enhanced synthesis of phenolics, flavonoids, terpenoids, and glucosinolates (GLSs) as adaptive responses. The review also highlights the molecular mechanisms underlying drought tolerance (DT), focusing on gene expression changes, signal transduction pathways, and the critical role of transcription factors (TFs). Abscisic acid (ABA) is pivotal in plant response and tolerance, regulating numerous genes through ABA signaling. TFs are central to genetic regulation under stress conditions, with several TFs, such as ABA-responsive element binding protein/factor (AREB/ABF), WRKY, NAC, and Nuclear factor‑Y (NF-Y), modulating gene expression in an ABA-dependent manner. Additionally, certain APETALA2/Ethylene response factors (AP2/ERF) and some NAC TFs govern gene expression independently of ABA. Furthermore, this study has highlighted the regulation of phytohormones and their interaction during drought events and the involvement of γ‑aminobutyric acid (GABA) signaling in DS management. The current study aims to provide a comprehensive understanding of ABA signaling in stomatal closure and the activation of TFs involved in regulating plant responses to DS. It also seeks to develop potential strategies to enhance plant tolerance to drought conditions.

Conserving mountain species under climate change is a major ecological concern. It is still unclear how potential climate change will affect mountain plant species. The present study investigates the potential distribution of Himalayan Boxwood (B. wallichiana) under current and future climate scenarios using an ensemble modelling approach. The results showed that the distribution of B. wallichiana is determined by mean annual precipitation, precipitation of the driest quarter and mean annual temperature. The model predicted that B. wallichiana has a narrow distributional range and is rarely found in the Himalaya under current climate. The future model predictions showed a considerable reduction in suitable habitats for B. wallichiana under all climate scenarios; more pronounced under PCP 8.5 for the year 2070. The model also revealed that currently suitable areas are likely to become unsuitable in the future and vice -versa. There is an urgent need to develop an appropriate policy for the conservation and management of B. wallichiana. The present study will contribute significantly to the formulation of appropriate policies for its conservation and management.

Oxidative stress, driven by reactive oxygen, nitrogen, and sulphur species (ROS, RNS, RSS), poses a significant threat to cellular integrity and human health. Generated during mitochondrial respiration, inflammation, UV exposure and pollution, these species damage cells and contribute to pathologies like cardiovascular issues, neurodegeneration, cancer, and metabolic syndromes. Lifestyle factors exert a substantial influence on oxidative stress levels, with mitochondria emerging as pivotal players in ROS generation and cellular equilibrium. Phytochemicals, abundant in plants, such as carotenoids, ascorbic acid, tocopherols and polyphenols, offer diverse antioxidant mechanisms. They scavenge free radicals, chelate metal ions, and modulate cellular signalling pathways to mitigate oxidative damage. Furthermore, plants thriving in high-altitude regions are adapted to extreme conditions, and synthesize secondary metabolites, like flavonoids and phenolic compounds in bulk quantities, which act to form a robust antioxidant defence against oxidative stress, including UV radiation and temperature fluctuations. These plants are promising sources for drug development, offering innovative strategies by which to manage oxidative stress-related ailments and enhance human health. Understanding and harnessing the antioxidant potential of phytochemicals from high-altitude plants represent crucial steps in combating oxidative stress-induced disorders and promoting overall wellbeing. This study offers a comprehensive summary of the production and physio-pathological aspects of lifestyle-induced oxidative stress disorders and explores the potential of phytochemicals as promising antioxidants. Additionally, it presents an appraisal of high-altitude medicinal plants as significant sources of antioxidants, highlighting their potential for drug development and the creation of innovative antioxidant therapeutic approaches.

Context Reproductive success in dioecious plant species may be limited by severe pollen limitation owing to their separate sexes and pollination barriers. Aims Dodonaea viscosa subsp. angustifolia (Jacq.) (Sapindaceae) is a dioecious and wind-pollinated species that has a long flowering period. This study sought to determine the relationship between its reproductive behaviour and pollen availability during different flowering phases. Methods Extra-stigmatic pollen germination and reproductive performance during different phases of flowering were investigated and correlated with pistil elongation under natural conditions. Results The species offers whole stigmatic and stylar surfaces for pollen to land and germinate under natural conditions. During pollen-limiting conditions, the length of the pistil increases significantly to enhance pollen capture. Depending on where on the pistil pollen lands, the timing of pollen tube arrival at the ovary varies. Conclusions Dodonaea viscosa subsp. angustifolia is the first reported wild species in the Sapindaceae where pistil elongation is regulated by pollination conditions and extra-stigmatic pollen germination ensures reproduction during phases of differing pollen availability. Our results indicated that the flexibility of female function and reproductive behaviour in Dodonaea viscosa subsp. angustifolia adds to the evolutionary possibilities to overcome pollination constraints. Implications This unique strategy for increasing female fitness through pollen presentation in D. viscosa subsp. angustifolia could be explored in other subspecies of D. viscosa on the Australian subcontinent.

Mountain ecosystems influence species distribution by offering climatic variables intertwined with rising altitude. These climatic factors determine species phenology and niche width. Although the distributional patterns of some prominent insect groups in relation to altitude have been determined, the environmental preferences along the altitudinal range that differentially influence the pollination of specific plant species are unknown. Here we assess how the composition and abundance of pollinator fauna of the important medicinal plant Berberis lycium Royle (Berberidaceae) differ across five distinct altitudinal gradients (800-2200 m) in the Pir-Panjal mountain range in the northwestern part of the Indian Himalayas. We monitored insect pollinators of major groups (bees, butterflies, wasps, flies) over two consecutive flowering seasons during 2022-2023. In total, 39 insect species belonging to five orders and 17 families were observed visiting the plant species during the flowering period across the altitudinal gradient. The results of the linear regression model depict that all four pollination indices show a negative correlation with increasing altitude in foraging activities when all the data are pooled together. However, only foraging speed (FS) and index of visiting rate (IVR) were statistically significant. In the case of individual orders, only Lepidoptera exhibited a notable relation to altitude. However, asynchrony in foraging activities among other pollinator groups has been reported along altitudinal gradients. The reproductive output (fruit and seed production) shows a significant negative correlation with increasing altitude. We concluded that while altitude influences species distribution, it also differentially shapes plant-pollinator interactions, pollinator activities, and reproductive output. This work is of great significance in order to monitor plant-pollinator interactions, which are essential component of biodiversity rich but fragile mountain ecosystem. ### Competing Interest Statement The authors have declared no competing interest.

Olea ferruginea, a wild widely distributed species of the olive lineage, exhibits andromonoecy with protogynous perfect (hermaphrodite) and staminate (male) flowers. The present communication elucidates the differential performance of staminate and perfect flowers in reproductive assurance. The flowering chronology and function of perfect and staminate flowers in the breeding system were investigated to understand the breeding behavior of the species. In terms of breeding behavior, pollen from staminate and perfect flowers was used to establish their role in breeding system under different pollination treatments. Results indicate higher pollen viability, germination, and pollen tube growth of staminate than perfect flowers. Results from pollen counts and morphological investigations show slight differences in both types. The species is self-incompatible which is indicated by pollen germination, the average number of pollen tubes in the style, and the fruit set following self and cross-pollination. Hand cross-pollination improves natural fruit set, especially when pollen from staminate flowers was used. Although the relict role of attracting pollinators cannot be fully discarded, it is suggested that the main benefit supplied by the formation of staminate flowers in olives is to promote male fitness by raising pollen output at the population level.

Plant reproduction and climate are strongly correlated with each other in various aspects, as climate change is the main cause of biotic and abiotic stresses, which adversely affect plant reproductive biology. We conducted a field experiment over two consecutive years to record how warming temperature affects flowering anthesis within inflorescence and the relationship between the flowers with anther dehiscence before anthesis (ADBA) and fruit set with maximum and minimum temperature in Olea ferruginea. The results of linear regression analysis show that there is increase in the rate of ADBA by 0.846 times per unit increase in maximum temperature and 0.789% per unit increase in minimum temperature. Also, there is decrease in fruit set by 0.933% and 0.743% when compared with maximum and minimum temperature respectively. The fruit set was found to be decreased by 0.81% when flowers undergo ADBA. Since the species is self-incompatible, cross pollination and pollen germination becomes impossible after anthesis due to the presence of self-pollen on the stigmatic surface. Our study results suggest that temperature fluctuations could affect rate of anthesis in flowering and reduce seed production under higher temperature. Thus, the impacts of climate extremes on plant life cycles may be as influential as gradual warming. In particular, it appears that throughout the anthropocene, temperature regime fluctuation had a bigger impact on flowering dynamics.

Olea ferruginea Royle is a wild Olive species commonly known as Indian or African Olive. In India, it is distributed in the Northwestern Himalaya, from Jammu and Kashmir to Uttarakhand, at an altitude of up to 2400 m amsl. The present study aims to determine the composition and community structure of Olea L. forests in Jammu and Kashmir, India. During the present study, 21 sites were surveyed from September 2019‒August 2021 and a total of 34 plant species belonging to 32 genera and 22 families were recorded. Fabaceae was the dominant family with five species followed by Moraceae (four species) and Fagaceae, Rosaceae, Malvaceae, Meliaceae, and Salicaceae (two species each). A total of 1333 individuals of trees were recorded including 362 individuals of Olea ferruginea. From the dendrogram resulting from Ward’s cluster analysis of 21 sites based on presence/absence data, five communities were identified on the basis of the Importance Value Index (IVI). The species richness and diversity indices showed great differences from community to community. The present study provides baseline information about the community structure and biological associations of Olea ferruginea.

Abstract A number of plants species believed to be wind pollinated (anemophilous) are actually pollinated both by wind as well as insects (ambophily) because all the traits of the two pollination modes have not evolved simultaneously. Comparatively few angiosperms exhibit ambophily and the significance of such a mechanism is not fully understood. During the present investigation, ambophily was studied in Olea ferruginea, over a period of three years. It is an andromonoecious and out-crossing tree species of the olive complex. The species is predominately pollinated by wind because it shows fruit set even when the insect pollinators were excluded. Moreover, a substantial amount of pollen was observed on the body parts of insect visitors and, comparatively, fruit set was enhanced in open-pollinated flowers receiving insect visitors. Floral structure of O. ferruginea displays floral traits favoring both wind and insect pollination. Furthermore, the flowers of this species were found to be visited by 20 insect species classified under three orders with Diptera and Hymenoptera being the leading pollen foragers. The results of linear regression of the hanging slide experiment showed that the dispersal of pollen grains decreased significantly with increasing distance from the canopy and the pollen density increased significantly with an increase in insect activity. The present work provides an insight to understand the ecological and evolutionary adaptations and consequences of spatio-temporal variation in plant – pollinator interactions. It is, therefore, useful for addressing both basic and applied questions in community structure and function including the evolution of floral traits, reproductive assurance, and the development of optimal conservation strategies.

The Himalaya represents the most fragile ecosystem in the world, signifying its sensitivity towards global climate change. In the current scenario, the Himalaya needs to assess the climatic change trends at the regional level. To investigate climatic trends, we analysed the long-term (1980–2020) climatic data (maximum temperature—TMax and minimum temperature—TMin, precipitation, and relative humidity—RH) collected by the five different meteorological stations from Shiwalik to Pir Panjal mountain range of Jammu province. The nonparametric Mann–Kendall test was used to analyse the significance of climatic change trends for temperature, precipitation, and RH data on seasonal and annual scales, whereas the nonparametric Sen’s estimator of the slope was applied to quantify the magnitude of climatic change trends. The results reveal that the Jammu province region received 53

Studies from different parts of the world have generated pieces of evidence of climate change’s effects on plant phenology as indicators of global climate change. However, datasets or pieces of evidence are lacking for the majority of regions and species, including for the climate-sensitive Himalayan biodiversity hotspot. Realizing this gap in information, and the wide-ranging implications of such datasets, we integrated real-time field observations and long-term herbarium records to investigate the changes in the spring flowering phenology of Olea ferruginea Royle, commonly known as the Indian Olive, in response to the changing climate in the western Himalayas. We attempted to create phenological change model using the herbarium records and field observations after recording the current dates of flowering and overall temperature trends from the study area over the last four decades from the five regional meteorological observatories of the Jammu province managed by Indian Meteorological Department (IMD) in Jammu and Kashmir. When considering current flowering dates along with herbarium information (years 1878–2008) for O. ferruginea, our Generalized Additive Model (GAM) showed 15–21 days-early flowering over the last 100 years significantly (p < 0.01). Results of the Mann–Kendall test showed increasing trends of TMin for all seasons significantly (p < 0.05) for Jammu province whereas TMax was only for the spring season. The increasing TMin of spring, summer, and autumn seasons also influenced the flowering phenology of O. ferruginea significantly (p < 0.01). By demonstrating the integrated use of methodological tools for finding long-term phenological changes in response to climate change, this work bridges knowledge gaps in phenological research from the developing world in general and the Himalayas in particular.

Recent shifts in the spectrum of weeds documented throughout the globe in various agroecosystems were purportedly associated with climate change. Consequently, it called for a more thorough investigation of weed vulnerabilities and management in the changing climate scenarios. In the present investigation, we compile geographic occurrence data of an agricultural weed Asphodelus tenuifolius Cavan. using ecospat R package and BIOMOD ensemble forecasting to analyze the existing distribution and anticipate its possible distribution in the future under future climate change scenarios. Nine different algorithms were used and implemented in biomod2 package for modelling the current potential habitat distribution of the selected species across the study region and climatic data was downloaded from WorldClim database. Ensemble modeling showed high habitat suitability of the majority of central, northern, and western Indian regions under current climatic conditions. In addition, by gaining suitable habitats in the future the target species will undergo significant increase in range change. The potential range of habitat suitability for A. tenuifolius is likely to increase from 138.73% to 190.29% in terms of Committee Averaging and 721.42%–1508% in terms of Weighted Mean under Representative Concentration Pathways (RCP 4.5 and RCP 8.5). This ensemble modelling technique will help us to locate the places where its potential range could expand while simultaneously analysing the spatial distributions of field weeds, which may be important for the creation of cutting-edge weed control methods.

Elevational gradient, slope, and aspect offer a unique opportunity to explore the response of plant species under changing environmental conditions. The present study aimed to analyze the species diversity and distribution patterns with respect to altitude, aspect, and habitat types in the Kashmir Himalayas. Considering major aspects and habitats, a total of 123 representative sites were selected along the elevational gradients for the present study. The plant species composition of each selected site was studied by organized sampling following the standard ecological methods. During the present study, a total of 361 vascular plant species belonging to 71 families and 214 genera were identified in the study area. At the lower altitudes, the southern aspect and drier habitats showed the highest diversity. Moreover, a significant amount of compositional dissimilarity was observed between the studied aspects, habitats, and elevation belts and was mainly due to species turnover rather than the nestedness component. Further, among the studied variables, altitude was the most important contributing variable, explaining the greatest variation in the species composition. The paired effects of altitude and habitat explained the maximum variation in plant species composition. It may be concluded that floristic diversity should be studied not only with reference to elevational gradients but should also include aspects and habitats. The current study will act as a reference in this direction. A similar study must be replicated in other parts of the Himalayan region in the future to improve our understanding of the distribution and preferences of plant species in mountainous zones. This, in turn, will be immensely helpful in the conservation and sustainable utilization of resources in these ecologically fragile regions.

Epigenetics are the heritable changes in gene expression patterns which occur without altering DNA sequence. These changes are reversible and do not change the sequence of the DNA but can alter the way in which the DNA sequences are read. Epigenetic modifications are induced by DNA methylation, histone modification, and RNA-mediated mechanisms which alter the gene expression, primarily at the transcriptional level. Such alterations do control genome activity through transcriptional silencing of transposable elements thereby contributing toward genome stability. Plants being sessile in nature are highly susceptible to the extremes of changing environmental conditions. This increases the likelihood of epigenetic modifications within the composite network of genes that affect the developmental changes of a plant species. Genetic and epigenetic reprogramming enhances the growth and development, imparts phenotypic plasticity, and also ensures flowering under stress conditions without changing the genotype for several generations. Epigenetic modifications hold an immense significance during the development of male and female gametophytes, fertilization, embryogenesis, fruit formation, and seed germination. In this review, we focus on the mechanism of epigenetic modifications and their dynamic role in maintaining the genomic integrity during plant development and reproduction.

In the era of anthropocene, global warming tends to alter the distribution range of the plant species. Highly fragile to such changes are the species that are endemic, inhabit higher elevations and show narrow distribution ranges. Predicting and plotting the appropriate suitable habitats and keeping knowledge of how climate change will affect future distribution become imperative for designing effective conservation strategies. In the current study we have used BIOMOD ensemble forecasting to study the current and predict the future potential distribution of Dactylorhiza hatagirea and Rheum webbianum and describe their niche dynamics in Himalayan biodiversity hotspots under climate change scenarios using ecospat R package. Results reveal sufficient internal evaluation metrics with area under curve (AUC) and true skill statistic (TSS) values greater than 0.8 i.e. 0.93 and 0.98 and 0.82 and 0.90 for D. hatageria and R. webbianum respectively, which signifies robustness of the model. Among different bioclimatic variables, bio_1, bio_3, bio_8, bio_14 and bio_15 were the most influential, showing greater impact on the potential distribution of these plant species. Range change analysis showed that both the studied species will show significant contraction of their suitable habitats under future climatic scenarios. Representative Concentration Pathway (RCP) 8.5 for the year 2070, indicate that the suitable habitats could be reduced by about 51.41% and 70.57% for D. hatagirea and R. webbianum respectively. The results of the niche comparisons between the current and future climatic scenarios showed moderate level of niche overlap for all the pairs with D. hatageria showing 61% overlap for current vs. RCP4.5 2050 and R. webbianum reflects 68% overlap for current vs. RCP4.5 2050. Furthermore, the PCA analysis revealed that climatic conditions for both the species vary significantly between current and future scenarios. The similarity and equivalence test showed that the niche between present and future climate change scenarios is comparable but not identical. From the current study we concluded that the influence of climate change on the habitat distribution of these plant species in the Himalayan biodiversity hotspots can be considered very severe. Drastic reduction in overall habitat suitability poses a high risk of species extinction and thereby threatens to alter the functions and services of these fragile ecosystems. Present results can be used by conservationists for mitigating the biodiversity decline and exploring undocumented populations on one hand and by policymakers in implementing the policy of conservation of species by launching species recovery programmes in future on the other. The outcomes of this study can contribute substantially to understand the consequences of climate change in the Himalayan biodiversity hotspots.

The structure and distribution patterns of Himalayan vegetation are poorly explored, and research on species composition along the elevation gradient in these mountain ranges is still deficient. The current study was undertaken to analyze the variation and pattern of plant species composition along a vertical gradient in northwestern Himalaya, India. A total of 18 sites were selected along an elevation gradient ranging from 2200 to 3900 m asl positioned at an interval of 100 m. The Renyi diversity profile, non-metric multidimensional scaling based on the Bray–Curtis dissimilarity metric and beta diversity components among the elevation belts were calculated. Furthermore, to study the influence of altitude on species richness and diversity, a generalized additive model was created. Two hundred and ten plant species representing 66 families and 147 genera were recorded. The Renyi diversity profiles show that the lower and mid-altitudes had rich species diversity. The results of the non-metric multidimensional scaling analysis show a considerable variation in the total plant species composition among the studied elevation belts. The observed multiple-site Sorensen dissimilarity index across the studied elevation belts was very high. The contribution of species replacement or the turnover component to the observed dissimilarity was much higher than the nestedness component. Furthermore, the herbaceous and tree richness showed a significant decrease with increase in elevation; however, the richness of shrubs showed a bimodal pattern. The present study increases our understanding of the trends and patterns of species richness along the vertical gradient in the Himalayan region.

Abstract Planning agricultural practises relies entirely on the timely prediction of rainfall based on data analysis. Early forecasting aids in the preparation of disaster management plans in high-risk locations in the event of predicted severe or limited rainfall. In this study, we analyzed the trends of precipitation and climatic variability for of Jammu region from 1925-2020. The non-parametric Mann-Kendall test was used to analyze the significance of trends in precipitation data on monthly seasonal and annual scales, whereas the non-parametric Sens’s estimator of the slope was used to quantify the magnitude of climatic trends. The results revealed that the Jammu region shows a statistically significant positive (p <0.005) for annual mean precipitation. In annual trend magnitude, the Jammu region showed a statistically significant increasing trend of 0.5260079 mm a-1 for the observed 95 years' climatic time series. The seasonal trends of precipitation statistics exhibit statistically significant positive trends over the observed time series in the case of the summer season only. Further, a significant precipitation increase of 1.484841 mm-1 was observed for the summer season only. The results of Pettit’s test for detecting annual change points for precipitation show a statistically significant change in the years 1988, 1951, and 1985 and seasonally in the year 1993 for the summer season only. Further, the results of the Mann–Kendall test for detecting monthly trends in the precipitation variables for the 95 years of observed climatic time series exhibit a statistically significant increasing trend for the months of May, June, August, and November. The results of this study are extremely useful in many sectors including agriculture, water resources, and most notably climatology studies in most striking aspects of developmental planning in recent times.

An increase in atmospheric greenhouse gases necessitates the use of species distribution models (SDMs) in modeling suitable habitats and projecting the impact of climate change on the future range shifts of the species. The present study is based on the BIOMOD ensemble approach to map the currently suitable habitats and predict the impact of climate change on the niche shift of Valeriana wallichii. We also studied its niche dynamics using the ecospat package in R software. Values of the area under curve (AUC) and true skill statistics (TSS) were highly significant (>0.9), which shows that the model has run better. From 19 different bioclimatic variables, only 8 were retained after correlation, among which bio_17 (precipitation of driest quarter), bio_1 (annual mean temperature), and bio_12 (annual mean precipitation) received the highest gain. Under future climate change, the suitable habitats will be significantly contracted by −94% (under representative concentration pathway RCP 8.5 for 2070) and −80.22% (under RCP 8.5 for 2050). There is a slight increase in habitat suitability by +16.69% (RCP 4.5 for 2050) and +8.9% (RCP 8.5 for 2050) under future climate change scenarios. The equivalency and similarity tests of niche dynamics show that the habitat suitability for current and future climatic scenarios is comparable but not identical. Principal Component Analysis (PCA) analysis shows that climatic conditions will be severely affected between current and future scenarios. From this study, we conclude that the habitats of Valeriana wallichii are highly vulnerable to climate shifts. This study can be used to alleviate the threat to this plant by documenting the unexplored populations, restoring the degraded habitats through rewilding, and launching species recovery plans in the natural habitats.

In the changing climatic conditions, species distribution modelling is considered as a key strategy to estimate the probable influence of climatic variabilities on the habitat ranges of any species. The present study explores the potential distribution of Aconitum heterophyllum under current and future climatic scenarios. The results unfold that the distribution of this endemic species is governed significantly by bio12, i.e., Annual Precipitation. Ensemble modelling predicted that higher altitudes of Jammu, Kashmir and Ladakh are suitable habitats for A. heterophyllum. However, the future climatic modelling revealed that there will be a significant decrease in the suitable habitats for A. heterophyllum. Most of the shrinkage of habitats is predicted to occur within the time period of 2050, which seriously challenges their survival. The present study recommends an urgent need to frame a pertinent conservation and management policy for Aconitum heterophyllum and will act as a framework for planning such a policy.

Climate change is one of the significant factors influencing global species redistribution. As a result, a better understanding of the species' possible range change in future climate conditions is needed. Therefore, this study compiles global geographic occurrence data of a wild olive sub-species, Olea europaea subsp. cuspidate, and projected potential distribution models in current and future climate scenarios. This study using ensemble modeling predicted that the species will undergo a significant decrease in habitat suitability under future climatic conditions with a contraction ranging from ca. 41 and 42% under RCP4.5 2050 and to about 56 and 61% under RCP8.5 2070 for committee averaging and weighted mean, respectively. More specifically, there will be a decrease in habitat suitability in regions of the southeastern part of the United States in North America; coastal regions in South America; coastal regions in the majority of eastern Africa; coastal parts of Spain, France, Italy, and Greece in Europe; coastal parts of Yemen and Saudi Arabia; the southeastern parts of Pakistan and the southern part of China in Asia; and southwestern and eastern parts of Australia when compared to current habitat suitability. The results of this ensemble modeling could be extremely valuable in identifying cultivation hotspots for the effective restoration and protection of this olive lineage under future climatic conditions.