ABSTRACT In the UK, large numbers of pheasants (Phasianus colchicus) are released into woodlands annually for recreational hunting. Post‐release, they are managed to keep them in areas where they can be shot from October to January. At sites with high pheasant densities, they can negatively impact local flora and fauna through predation, trampling, and altering soil nutrients via defecation. The extent of these effects beyond release pen sites is unclear. This study investigated at what distance from release sites a suite of ecological effects of released gamebirds might be detected. We surveyed along 1 km transects from release pens at 20 shoots in Ancient Semi‐Natural Woodlands (ASNW) in England and Wales. We expected higher soil nutrients, fewer characteristic woodland plants, more nitrogen‐tolerant species, more damage to dead wood, and fewer woody seedlings near pens, with effects decreasing with distance. We found that pheasant numbers encountered declined with distance and sites closer to pens had fewer seedlings/saplings, lower vascular plant richness and less decayed wood (although this last result may be an artefact of the study design). These effects did not extend beyond 500 m. Contrary to expectations, soil nutrients, bare ground, and ancient woodland indicator species showed no consistent change with distance, and nitrogen‐loving species and weeds were more common further from pens. We conclude that, in areas beyond release pens in ASNW in lowland England and Wales, where pheasants are not deliberately enticed, any direct negative effects on plant communities, soil nutrients and ground cover do not extend further than 500 m from the point of release. The current licensing of gamebird releases in England aims to protect designated woodlands by controlling releases occurring within a 500 m buffer around protected areas. Our results suggest that this buffer size appears to be appropriate to contain these effects of released gamebirds.

Abstract The red fox, Vulpes vulpes , is abundant in England and can exert limiting effects on their avian and mammalian prey. Large-scale gamebird releases in the UK may be sustaining high predator numbers, leading to greater predation on other prey species, especially once gamebird stocks are depleted. However, little is known how gamebird release affects the broader diet of the red fox. Here we used DNA metabarcoding to assess the diet of foxes from 18 agricultural estates in Southern England, 10 of which released large numbers of gamebirds (red-legged partridges – Alectoris rufus , and common pheasants – Phasianus colchicus ) and 8 which did not. Scats were collected over one year, allowing for seasonal investigation of the foxes’ diet. We investigated the vertebrate species consumed and compared the non-gamebird dietary diversity and composition between release and non-release estates and across seasons. The field vole ( Microtus agrestis ) was found to be the most frequently predated species overall. Brown hares and field voles were detected more on release sites, while bank voles and dog faeces were detected more on non-release sites. We found little evidence that foxes predate ground nesting birds or other species of concern. The dietary diversity was significantly lower on estates that released gamebirds, and this difference was most notable during the post-shoot, spring months (February to April). On both estate types, diversity was highest in the summer months. The altered predatory behaviour due to gamebird release is likely to affect the populations of the non-game prey of the red fox and this should be considered when designing policies regarding gamebird management and biodiversity conservation. Alternative predation combined with predator control may be reducing predation pressure on non-game prey where gamebirds are released, but high fox density elsewhere likely results in higher predation pressure on a wide range of species.

ABSTRACT Large-scale annual releases of pheasants Phasianus colchicus and their subsequent management for recreational shooting create various ecological impacts in the UK. While effects at release sites are fairly well understood, dispersing birds may influence areas farther away. If they enter ecologically important but sensitive protected areas (PAs), any negative impacts could be especially harmful. Using tracking data, from 766 birds across 10 sites, we estimated survival and dispersal of released pheasants and applied these patterns to gamebird release records near English PAs to gauge intrusion risk. Of 2,885 registered release sites, just over half lay within 2 km of a PA. A large number of shoots release relatively few birds while a small number release many birds. Thus, numbers expected to enter a particular PA likely depend both on the size of releases and proximity to the PA. We estimate that, at a national level, a maximum of between 525,000 and 784,000 pheasants might be found within PAs very soon after release, representing around 1.7% of all the pheasants released annually. This number declines over the months after release until in February, we estimate that there are between 131,000 and 196,000 pheasants (0.4% of the total release) might be found within PAs. The critical metric by which ecological damage might occur is their density within PAs. Mean densities soon after release averaged 12.0 birds/ha in PAs within 250 m of release sites. This density declined markedly both in time (as birds died) and space (as they moved further from the pen as potential areas increased). By November, densities in PAs 500-1000m from release sites peaked at 0.5 birds/ha, falling to 0.16 birds/ha in February. These estimated densities are around two orders of magnitude lower than those known to cause strong, lasting impacts within release pens. The results are subject to assumptions about movement behaviour, game management and bias in registration. Despite these constraints, considerable local variation exists, with a minority of high-volume release sites very near PAs posing the greatest potential ecological risk.

CapsuleDuring autumn and winter a small proportion of summer-released Common Pheasants or Red-legged Partridges disperse beyond the release/shooting area and occupy nearby land in the following spring at densities of between 5 and 60 individuals per km2.AimsTo compare densities of released Common Pheasants and Red-legged Partridges on land used for shooting and on non-release, non-shooting land nearby.MethodReleased birds were surveyed along walked transects on ten large release sites in southern England, and on eight non-release sites within 2 km of a release site, every three weeks for 19 months and over two summers. Proven spring count methods were used to adjust long-term survey data to estimate approximate densities.ResultsIn early spring, after shooting had ended, there were 100-450 Common Pheasants per km2 at release sites, and 50-250 Red-legged Partridges. In early summer, before the next release, there were up to 100 Common Pheasants and 70 Red-legged Partridges per km2. Non-release sites always had significantly fewer birds: in early spring there were 20-60 Common Pheasants and 5-25 Red-legged Partridges per km2; in summer there were fewer than 20 Common Pheasants and ten Red-legged Partridges per km2 on all but one non-release site. Following the next summer release and initial dispersal, we documented a circa 50-fold increase in densities, with 300-2,500 Common Pheasants or Red-legged Partridges per km2 on shooting land in autumn. On the non-release sites, a modest and delayed response to dispersal from release sites involved a 50-100% density increase over winter.ConclusionsOur proportion and density estimates can help conservationists to understand the potential ecological effects of releasing Common Pheasants and Red-legged Partridges for shooting, and support modelling work that can predict distributions of released birds in different scenarios to help guide release policies.

The cause of recent population declines in some farmland/hedgerow breeding bird species in the UK is related to a lack of cover and food resources in winter. In improved grassland areas some of those declines have been particularly acute and some have been shown to be related to the availability of grass and weed seed in winter. The provision of seed‐bearing crops as part of AES options has been shown to benefit some of these birds. Game crop plots sown on shooting estates for holding and driving gamebirds in autumn and winter have been shown to hold relatively high densities of farmland and wood‐edge birds during the winter. We studied breeding songbirds in hedges in a primarily improved grassland area in the SW of England where there are some large shooting estates that sow relatively large game crop plots (1–5 ha) in the landscape. In this study we found that otherwise similar hedges in terms of size and density near to those winter game crop plots, had between 1.5 and 2 times as many breeding resident songbirds per unit length the following spring compared to hedges further away from game crop plots. This was despite game management in these plots being wound down during February and in many cases, the crops themselves being removed by mid‐March. Hedges within approximately 350 m from game crop plots had more breeding birds. We discuss possible mechanisms and suggest that some passerines preferentially establish breeding territories in hedges near to game crops in late winter. We suggest how to distribute game crop plots to maximise any benefit in an improved grassland landscape.

This expert opinion discusses the mechanisms by which climate and management could alter tick and game host (animal) densities and contact rates, as well as considering the likely relative importance of climate change and game (animal) management in affecting the distribution of ticks in Europe.

Owing to the long-term decline of wild gamebird populations and the advances on poultry production, rearing has become increasingly popular for put-and-take shooting rather than for re-establishing wild populations. Although hunters may prefer shooting wild gamebirds, reared ones ensure hunting bags at a reasonable cost; hence the majority of gamebird shooting in Europe, either commercial or non-commercial, rely on reared birds. This chapter reviews current knowledge on gamebird rearing and releasing, with a special focus on red-legged partridges. It is estimated that around 14 million red-legged partridges are reared for shooting in Europe annually, being the vast majority of them reared under commercial methods. Research confirms that commercial rearing and releasing cause a long list of detrimental effects on reared birds, their wild counterparts and other wildlife. However, non-commercial rearing and releasing for re-establishment attempts may be helpful for conservation purposes; reared birds in general will fly well in front of guns, and a small proportion may breed if given the chance. To ensure the fate of wild gamebirds' populations and the feasibility of rearing, we suggest changing from large-scale releasing for shooting into releasing for conservation. To do so, quality labels to certify estates producing wild red-legged partridges should be launched, and wild gamebird management should get funding (private and public) and recognition. We suggest ways to improve commercial rearing and provide guidelines for rearing and releasing for re-establishment purposes.

This review examines 128 items of primary and other literature to provide an insight into current knowledge of the effects of pheasant and red-legged partridge releasing and associated management for shooting on habitats and wildlife in the UK. It summarizes key findings and uses them to define sub-topic sections for which the effects are classified as positive, neutral or negative. This forms the basis of a numerical synthesis of effects and some overall conclusions. Fifty-four directly related studies were identified, which defined 25 sub-topics or effects. A mix of positive, neutral and negative ecological consequences of releasing are described, for which the corresponding number of sub-topics approximately balance each other. Positive effects are usually a consequence of gamebird management activities, most negative effects are caused by the released birds themselves. The different spatial scales at which effects are likely to operate are identified, for example effects on generalist predators or of gamecrops occur at the landscape scale, while many habitat effects have a local impact. Some local negative effects have relatively straightforward management solutions for example, by identifying and avoiding especially sensitive sites when locating release pens. The synthesis identifies seven negative effects associated with the increasing scale of releasing. Several positive effects are linked to economic considerations and are more likely to have greater impact at larger shoots. Pheasants released into woodland have more direct local effects than partridge releases on farmland. The framework of sub-sections could be used as the basis for a more complex synthesis or weighted analysis for a particular set of ecological priorities. The review findings should be interpreted as representing a median type of shoot in terms of size and adherence to good practice over recent decades. They increase the awareness of potential conflicts, highlighting the need for best practice and what factors to consider for mitigation.

The release of large numbers of juvenile pheasants into open-topped release pens in woodlands is a common part of game management in the UK. Previous research has shown this practice modifies the soil conditions and ground flora community of these release pens. However, it is not currently known if and how these changes to soil and ground flora reverse once the pens are no longer used. We compared the soil chemistry, ground flora structure and community composition of disused release pen sites in ancient semi-natural woodlands with paired control sites. Some of the changes seen within release pens in active use persisted in disused pens; soil fertility and cover of species that prefer fertile soils were higher in disused pens, whereas winter green perennials, richness of species of ancient semi-natural woodland and overall species richness were lower. Total species richness and richness of ancient semi-natural woodland plants showed signs of recovery in pens that had been disused for longer than ten years, but this recovery only occurred in pens where ≤1000 pheasants/ha had been released. Pheasant release pens are sometimes relocated within woodland to reduce disease incidence but, as the flora within disused pens does not recover quickly, this practice may cause cumulative habitat damage. We recommend that release pen relocation should be minimised and suggest other management strategies that could reduce the need to relocate pens and increase the floral recovery in disused pens, such as reducing the density of pheasants released.

The recent decline in farmland songbirds in the UK has coincided with increases in the populations of many nest predators. However, studies which have removed nest predators and monitored the response of prey populations have found mixed results. One explanation for this ambiguity is that, within species, predators differ in how likely they are to predate nests and only the removal of particularly predatory individuals will improve the breeding success of prey populations. Predators could differ in the extent to which they take nests due to variation in the local environment and/or variation within the predator population, e.g. differences in breeding status. Additional to these broad factors, certain individuals may specialise on particular prey. We placed 460 artificial nests in a systematically balanced design in UK farmland to analyse these sources of variation in predation. Magpies (Pica pica) were the most common predators of our artificial nests and the vulnerability of our nests to magpie predation varied according to magpie breeding status (predation was higher inside breeding magpies' territories), but this effect varied temporally. More nests were predated inside of magpie territories late in the season, when magpies had dependent fledglings. More specifically, some nest locations were especially vulnerable independent of both magpie breeding status and time in breeding season. These nests may have been disproportionately predated by specific, particularly predatory, territorial magpies. Habitat management and/or predator removal may benefit songbird populations if targeted towards reducing the effect of particular individuals identified as more likely to predate songbird nests.

We present data accumulated over the last 25 years on predation of radio-tracked released pheasants. In studies of birds during the autumn/winter at six pheasant shoots with high-density releases managed by full-time gamekeepers, predation of released pheasants by foxes before the shooting season began (July–September) averaged 19.2 ± 4.0% per site, and during the shooting season (October 1st–February 1st), a further 15.9 ± 1.9% were predated. The range in 3-year average predation rates between sites before shooting began was 8.6 to 42.4%. At seven different sites during the spring and summer, between 20 and 71% of released or wild hens that survived the shooting season were predated, mainly by foxes, between mid-March and mid-July. Predation was significantly higher at sites with low-level predator control (59 ± 4.7%) compared to those with high-level control (30 ± 5.3%). At three of the four sites with low predator control between 5 and 22% of nest failures were caused by incubating hens being predated by foxes. Our data quantify for the first time highly variable predation rates of released pheasants before and during the winter shooting season which we suggest was influenced by a range of site and management factors. During the spring and summer, our data provide evidence that predation of adult hen pheasants as well as nest predation can suppress breeding success and that predator control can reduce these losses.

The role of predation by corvids on the breeding output of songbirds is unclear. Using a randomised‐pair design, we measured how nest success of hedgerow‐nesting passerines responded to the experimental removal of carrion crows and magpies. We worked in southern England at 32 paired sites around 4 km2 each, one with and one without best‐practice corvid control, studying four different pairs per year for four years 2011–2014. We counted corvids, and using songbird territory mapping and fledged brood counts without finding nests along transects, we estimated nest success as a brood/ territory ratio for the community of songbirds in 4 km of hedgerow at each site. Crows and magpies were still present at most removal sites but numbers were half as high as at paired non‐removal sites. Eighteen songbird species were frequently encountered at most sites with on average (±1 SD) 102 ± 30 territories per site. Using a generalised linear mixed model analysis the songbird community as a whole bred less well in treatment sites without corvid removal and in years with more rainfall. Nest success was down by 10% in non‐removal sites on average relative to removal sites over the four years. Excluding 2012 data because of exceptionally high spring rainfall that year, nest success was down 16% in the non‐removal sites on average in the other three years. For open‐cup nesting species as a group there was no difference in nest success between site types. Our data on hole nesters suggest that they were affected by treatment and contributed to our overall result. For species whose numbers are regulated through territoriality, nest‐site or habitat availability, spring abundance is unlikely to be affected by a 15% increase in breeding output. For species limited by nest success, it may be more important.

Subtle variations in early rearing environment influence morphological, cognitive and behavioural processes that together impact on adult fitness. We manipulated habitat complexity experienced by young pheasants (Phasianus colchicus) in their first seven weeks, adding a third accessible dimension by placing elevated perches in their rearing pens mimicking natural variation in habitat complexity. This simple manipulation provoked an interrelated suite of morphological, cognitive and behavioural changes, culminating in decreased wild mortality of birds from complex habitats compared with controls. Three mechanisms contribute to this: Pheasants reared with perches had a morphology which could enable them to fly to the higher branches and cope with prolonged roosting. They had a higher propensity to roost off the ground at night in the wild. More generally, these birds had more accurate spatial memory. Consequently, birds were at a reduced risk of terrestrial predation. The fitness consequences of variation in early rearing on behavioural development are rarely studied in the wild but we show that this is necessary because the effects can be broad ranging and not simple, depending on a complex interplay of behavioural, cognitive and morphological elements, even when effects that the treatments provoke are relatively short term and plastic.

Suggestions that novel, non-food, dedicated biomass crops used to produce bioenergy may provide opportunities to diversify and reinstate biodiversity in intensively managed farmland have not yet been fully tested at the landscape scale. Using two of the largest, currently available landscape-scale biodiversity data sets from arable and biomass bioenergy crops, we take a taxonomic and functional trait approach to quantify and contrast the consequences for biodiversity indicators of adopting dedicated biomass crops on land previously cultivated under annual, rotational arable cropping. The abundance and community compositions of biodiversity indicators in fields of break and cereal crops changed when planted with the dedicated biomass crops, miscanthus and short rotation coppiced (SRC) willow. Weed biomass was consistently greater in the two dedicated biomass crops than in cereals, and invertebrate abundance was similarly consistently higher than in break crops. Using canonical variates analysis, we identified distinct plant and invertebrate taxa and trait-based communities in miscanthus and SRC willows, whereas break and cereal crops tended to form a single, composite community. Seedbanks were shown to reflect the longer term effects of crop management. Our study suggests that miscanthus and SRC willows, and the management associated with perennial cropping, would support significant amounts of biodiversity when compared with annual arable crops. We recommend the strategic planting of these perennial, dedicated biomass crops in arable farmland to increase landscape heterogeneity and enhance ecosystem function, and simultaneously work towards striking a balance between energy and food security.

Behavioural and physiological deficiencies are major reasons why reintroduction programmes suffer from high mortality when captive animals are used. Mitigation of these deficiencies is essential for successful reintroduction programmes. Our study manipulated early developmental diet to better replicate foraging behaviour in the wild. Over 2 years, we hand-reared 1800 pheasants (Phasianus colchicus), from 1 day old, for 7 weeks under different dietary conditions. In year one, 900 pheasants were divided into three groups and reared with (i) commercial chick crumb, (ii) crumb plus 1% live mealworm or (iii) crumb plus 5% mixed seed and fruit. In year two, a further 900 pheasants were divided into two groups and reared with (i) commercial chick crumb or (ii) crumb plus a combination of 1% mealworm and 5% mixed seed and fruit. In both years, the commercial chick crumb acted as a control treatment, whilst those with live prey and mixed seeds and fruits mimicking a more naturalistic diet. After 7 weeks reared on these diets, pheasants were released into the wild. Postrelease survival was improved with exposure to more naturalistic diets prior to release. We identified four mechanisms to explain this. Pheasants reared with more naturalistic diets (i) foraged for less time and had a higher likelihood of performing vigilance behaviours, (ii) were quicker at handling live prey items, (iii) were less reliant on supplementary feed which could be withdrawn and (iv) developed different gut morphologies. These mechanisms allowed the pheasants to (i) reduce the risk of predation by reducing exposure time whilst foraging and allowing more time to be vigilant; (ii) be better at handling and discriminating natural food items and not be solely reliant on supplementary feed; and (iii) have a better gut system to cope with the natural forage after the cessation of supplementary feeding in the spring. Learning food discrimination, preference and handling skills by the provision of a more naturalistic diet is essential prior to the release of pheasants in a reintroduction programme. Subsequent diet, foraging behaviour, gut morphology and digestive capabilities all work together as one nutritional complex. Simple manipulations during early development can influence these characteristics to better prepare an individual for survival upon release.

The management of pheasants (Phasianus colchicus) in pens and their subsequent release within woodland for game shoots is widely practised in Britain. With the exception of ground flora and songbirds, the impacts on other taxa have not been well documented. We investigated the effects of pheasants on invertebrate abundance and community composition, using pitfall trapping. We compared release pens with control plots located in the same woods and in woods where no pheasant releasing had occurred for at least 25years. Conditions for invertebrates within release pens were altered, with more annual plants and disturbance-tolerant perennials and a reduced leaf litter layer. No major differences in invertebrate abundance, or Carabidae or Staphylinidae richness, were found in spring at either the pen scale or the wood scale. However, pheasant release pens resulted in significant changes in the species composition of Carabidae, with shifts towards species typical of arable fields and grassland. Carabid species active in spring and those that are very large (>17.0mm) declined at pheasant release densities higher than 1000birds/ha. Both effects are likely to be due to predation by pheasants at the peak of release in July–August, operating separately on larvae and adults respectively. There was an overall increase in the abundance of detritivores, including Diplopoda, Oniscoidea, Gastropoda (snails), at higher release densities. Mean release density in our study was 1489±126birds/ha (range 174–3409, n=37 pens) and we suggest that detrimental effects on specialist woodland invertebrates would be minimized if releasing was conducted at the recommended density of 700birds/ha.

A key aim of many European agri-environment schemes (AES) is to improve biodiversity on farmland. In recent years, several countries have been looking at long term trends in the spring adult population size of a target group of farmland birds as an indicator of this. The overall trend in these indicators is however not upwards. While this might suggest we need to look at the design and deployment of habitat management options within schemes, there is an increasing view that we also need to refine and improve our indicators, or the way we monitor them.Relating spring adult bird population size to AES options is problematic not least because of the time lag between the deployment of those designed to enhance bird breeding success in summer, and the spring surveys the following year. At the other end of the scale detailed studies of breeding success in farmland birds have practical/cost restraints. We argue that to understand the impact on farmland birds of particular summer options within AESs we need to be able to estimate the breeding success of local bird populations quickly and cheaply. This would enable us to relate particular AES options to the breeding performance of the birds actually using them.Complementing a previous study of woodland birds, we assess the likelihood of encountering fledged broods of hedgerow nesting bird species during transect surveys without finding nests, and then apply a simple mark-recapture analysis technique to provide an index of breeding success for those species. Following spring adult assessments, counts of fledged broods were undertaken four times a week during April, May, June and July, in four 2.5 km hedgerow transects, at four sites in southern England in 2010. Mean daily detection probabilities of fledged broods of 16 common hedgerow birds were calculated from these counts using the software Presence. For 15 out of these 16 species these detection probabilities were sufficiently high for a programme of fledged brood surveys, involving just two or three visits per week from mid-May to mid-July, to provide a useful estimate of breeding success.The survey technique and associated analyses make certain assumptions when providing estimates of breeding success and these are discussed. Little is known about initial dispersal in passerine fledglings and a study in hedgerows may be useful here. However our pilot study suggests that the method could have application as a relatively easily derived productivity index for hedgerow birds, and hence an additional method available to study the impact of certain AES options on indicator species, or for research studies. (C) 2015 Elsevier Ltd. All rights reserved.

Polyommatus bellargus is a priority species of butterfly in the UK as a result of its scarcity and the rate of population decline over the last few years. In the UK, the species is associated with chalk grassland on hot, south-facing slopes suitable for the growth of the food plant Hippocrepis comosa. Shooting game birds is a popular pastime in the UK. Over 40 million game birds, principally Phasianus colchicus and Alectoris rufa, are bred and released into the countryside each year for shooting interests. There is a concern that the release of such a large number of non-native birds has an adverse effect on native wildlife. A study was carried out over a period of 3 years out to examine whether there was any evidence that A. rufa released into chalk grassland habitat negatively affects populations of P. bellargus. A comparison was made between sites where large numbers of A. rufa were released versus sites where no, or few, birds were released. The study involved the construction of exclosures in these sites to allow an examination of the number of butterflies emerging from H. comosa when the birds were excluded versus when the birds had free range across the area. Where birds were present the on-site vegetation was shorter than where they were absent indicating that the birds were definitely influencing habitat structure. However, the evidence that A. rufa was negatively influencing the number of adult butterflies emerging was not strong, although there was a largely non-significant tendency for higher butterfly emergence when the birds were excluded or absent.

Capsule Summer bird densities in establishment-stage (2-3-year-old) Miscanthus were the same as in winter wheat fields for many species, but were higher for Lapwing, and for Reed Bunting and Blackbird late in the summer.Aims To compare abundances of farmland birds in establishment-stage Miscanthus biomass crops with those in crop types they are likely to replace (winter wheat and grassland) during the breeding season. In particular, to investigate whether such a rapid-growing, dense crop has lower abundances of field-nesting species.Methods Bird surveys were conducted in 51 Miscanthus fields and an equal number of grass or winter wheat control fields between May and July, in Lincolnshire and southwest England.Results Species richness was similar in Miscanthus to that in winter wheat and grass. Skylark densities were similar in Miscanthus and wheat, while Lapwing densities were higher in Miscanthus and grass than wheat. Miscanthus contained very high densities of Reed Buntings and Blackbirds late in the summer.Conclusion Densities of most bird species, including field-nesting species, were not lower in establishment-stage Miscanthus than in the crops it most commonly replaced. The Miscanthus surveyed was relatively weedy and patchy; further studies as crops age and knowledge of Miscanthus husbandry increases would be valuable.

Hedgerows are an integral part of the farmed landscape and are important habitats for many birds and other wildlife. Game managers recognize the importance of hedgerows for gamebirds, both in the nesting season for wild birds and as dispersal routes from woodland release pens for reared pheasants in late summer. Conservationists are interested in the potential impacts of pheasant releasing on wildlife and habitats. The aim of this study was to assess the impact of gamebird management on hedgerow structure and the density and connectedness of hedgerows at the landscape level by comparing sites with and without gamebird releasing. We surveyed 150 sites in southern and eastern England: 97 were on pheasant shoots while the others were on farms where there had been no pheasant releasing for at least 25 years. Using a combination of field survey, satellite imagery and GIS software, we determined hedgerow abundance and their structural characteristics. We also measured the extent of hedgerow connectedness. We found that the hedgerow structure was similar on farms with and without game shoots. However, hedgebanks were between 17% and 36% wider on game estates. Game estates also had between 10% and 65% more hedgerow per square kilometre than farms with no game shoot. We found no strong evidence that game management influenced bird numbers associated with individual hedgerows. We conclude that structural characteristics of hedgerows on sites with pheasant releasing do not differ from hedgerows on farms where there is no pheasant releasing undertaken. The tendency for wider hedgebanks and greater hedgerow abundance on game estates is likely due to the recognition of game estate managers of the value of hedgerows as habitats for gamebirds. The results of this study need to be considered within the context of other studies which have highlighted the potential negative effects of large pheasant releases on hedgerow vegetation.