The shell midden at Fårevejle was first visited by A.P. Madsen in 1894 and later partially excavated in 1896-97 by the National Museum of Denmark. In 2004-2005 the shell midden was revisited; two trenches were put through the central part of the midden. The shell midden is clearly stratified with Ertebølle layers at the bottom; the main shell component of these layers is oysters. These layers contain stone-lined fireplaces, bone, diagnostic pot sherds, and flint artefacts from the Late Mesolithic. The top of the midden is mainly composed of crushed cockles and mussels, mixed with layers of charcoal and burned rocks. These upper layers contain fireplaces and ash dumps, as well as bone, flint artefacts and potsherds from the EN Ib/EN II and MNA Ib phases of the Funnel Beaker Culture (TRB). The sparseness of residential materials (flint, sherds, bone) in the Mesolithic component suggests that this midden was not a base camp but rather visited for short periods of use.

Animal bones from archaeological contexts can reveal the interplay between past environments and human societies. Resource acquisition shaped many aspects of past societies and influenced the development of trade networks and migration. Fish have been a cornerstone of human subsistence for millennia, yet the rise of commercial fishing and trade was complex. Here, we synthesised a database of ∼1.9 million zooarchaeological fish records spanning 2000 years across Europe. Using machine-learning of catch compositions alongside fish thermal tolerances, we show that fisheries became less local over time, with homogenisation coinciding with Little Ice Age-associated cooling, a period of documented resource scarcity, concurring with growing trade. Moreover, increased proportions of marine taxa and more specialist marine fisheries were observed in the preceding Medieval Climate Anomaly, to sustain concurrent urban and population growth. Enhanced use of marine protein buffered food insecurity, whilst signalling the transition from localised to trans-regional trade networks. ### Competing Interest Statement The authors have declared no competing interest. European Research Council, https://ror.org/0472cxd90, 951649

Abstract Fish played a vital role in the development of the European Mesolithic; from their colonization of the continent during the Upper Palaeolithic to ongoing debates regarding their importance across the Neolithic transition, they are central to many discussions. Here, we focus on the European Mesolithic zooarchaeological fish dataset, in which we have collated >640,000 identified fish remains from 358 sites. We aim to document the development of fishing from the Early-through-Middle Holocene across the continent. While acknowledging certain limitations—for instance preservation and recovery biases, research histories, and language constraints—by assembling this comprehensive dataset it is possible to begin to evaluate how fishing developed during the Mesolithic. Importantly, it is possible to compare the trajectories evidenced from different areas. The transition from freshwater to marine-focused fisheries and the development of distinctive local traditions are highlights of the research.

New excavations of an enclosure-related site at Ginnerup on Djursland, Denmark, in 2020 resulted in the identification of several features containing depositions of marine shells. One of these, A4, is a natural depression with a fill comprised of four consecutively deposited layers, forming an undisturbed stratigraphy, dated by several 14C dates to between c. 3150 and 2950 BC. The oldest layer contained finds from phase MN A Ib of the Funnel Beaker culture, while the remaining three layers yielded finds from the latest Funnel Beaker culture on Djursland (MN A II/III, Ferslev style) with an upwardly increasing content of Pitted Ware culture elements, thereby allowing the emergence of this culture in Denmark to be followed for the first time. Preservation conditions for organic material were excellent due to a content of marine shells, mainly from oysters and mussels, in all layers. In this preliminary account, a survey of the material culture in the four layers is presented, together with 14C dates, zoological investigations of mammal and fish bones, isotope analyses (d13C, d15N and d34S) and aDNA analyses of mammal bones and examinations of plant macro-remains. The abundant bones of wild horses also hold a huge potential for zoological and genetic studies, the results of which can qualify the ongoing debate about the rewilding of horses in present-day Europe.

The spread of the Neolithic way of life from its centers of origins remains one of the central topics of archaeological research, with ongoing debates about the importance of economic, demographic, and cultural changes in the transition. The Southern Caucasus, while close to one area where agriculture emerged, has remained understudied regarding this spread. Here, information about the role of fish, a topic that has been almost completely neglected until now is presented. Fish remains are scarce in this region. Moreover, isotope analyses seem to indicate that freshwater fish were not an important food source. For the first time, fishbones have been found in larger quantities at the site of Aruchlo I from some layers in ditches. It is the largest assemblage of fish bones safely dated to the sixth millennium BC in the South Caucasus. The interpretation of these finds is not straightforward due to the lack of other comparable finds and the absence of fishing gear. Fishing appears to have been conducted in the waters close to the settlement. It is unclear if fishing was a year-round activity, although the way these bones were concentrated in different layers in the ditches suggests that this was not the case. We think that the bounteous catch of spawning fishes at certain times of the year can be linked to special social events like feasting, showing the importance of a food resource that is usually greatly underrepresented archaeologically. Introducing more precise recovery methods for animal remains at other excavations will hopefully refine our understanding.

Exploitation of aquatic resources, especially fishing, was a fundamental part of human subsistence during the Late Mesolithic Ertebolle culture (5400-4000/3900 cal BC) in Southern Scandinavia. In this pilot study we examine three cod otoliths from two Late Mesolithic locations in eastern Denmark to see whether local environmental conditions are reflected in the strontium ratios of the fish, to source the fish to either the Atlantic or the Baltic Sea and finally, to explore how fishing was conducted in relation to the settlements. We used laser ablation multi-collector inductively coupled plasma mass spectrometry for sequential sampling of the otoliths for strontium isotope analysis. All three otoliths yielded Sr-87/Sr-86 values that fall within the range of Baltic Sea water, thus indicating that cod caught by Mesolithic fishers in the Danish straits belonged to the Baltic Sea stock. Our results suggest that cod were not caught in waters immediately adjacent to the archaeological sites but rather came from the Kattegat. We could not detect any substantial change in habitat between juvenile and mature stages of the fishs lives. Our study shows the potential of isotopic analysis to address issues regarding the individual ecological history of fish and human fishing strategies.

This paper provides a new synthesis of perforated fish vertebrae in prehistoric and historic Europe, with a particular focus on Poland, within the broader context of Central and Eastern Europe. The earliest examples of such artefacts in Europe date from the Upper Palaeolithic, but compared to other 'beads' manufactured from animal bone, perforated fish vertebrae are rare. This paper examines the diachronic trends in the range of species that have been chosen for such objects, as well as their depositional contexts. Despite the wide range of freshwater and marine species exploited by people, only the vertebrae of a few species-especially pike and catfish-were selected for use as beads. There is a general shift from their deposition in funerary contexts in prehistoric European societies to their association as low-status objects associated with Christian private devotion in the post-conversion period. However, this may not reflect continuity in the use of fish vertebrae, with a shift in their symbolism after the conversion to Christianity, given the substantial chronological gaps in the archaeological record. This synthesis nonetheless provides a solid foundation for contextualising future archaeological finds of such artefacts.

Southern Scandinavian Mesolithic research has one of the longest traditions within archaeology, dating back to the 1820s and 1830s.However, a combination of site visibility and an emphasis on the Mesolithic-Neolithic transition has meant that research has primarily been directed towards the Late Mesolithic Ertebølle culture (c. 5400-4000 cal.BC) at the expense of the Early Mesolithic Maglemose culture (c. 9600-6400 cal.BC).Whilst fishing during the Ertebølle culture is well studied (Enghoff 2011; Ritchie 2010), fishing during the Early Mesolithic is rarely discussed in any detail.In this contribution we attempt to rectify this imbalance by collating all readily available data on fish remains and related technologies within the literature.Although our primary focus is the Early Mesolithic Maglemose culture of Southern Scandinavia, an area encompassing Denmark, Scania in Sweden and Schleswig-Holstein in Northern Germany, we draw on contemporaneous sites within the broader region to provide a more nuanced picture of the exploitation of this important resource, fish.

The Stone Age site Riņņukalns, Latvia, is the only well-stratified shell midden in the Eastern Baltic. In this paper, we present new interdisciplinary results concerning its dating, stratigraphy, features, and finds to shed light on the daily life of a fisher population prior to the introduction of domesticated animals. The undisturbed part of the midden consists of alternating layers of unburnt mussel shell, burnt mussel shell and fish bone, containing artefacts, some mammal and bird bones, and human burials. Two of them, an adult man and a baby, are discovered recently and date to the calibration plateau between 3350 and 3100 cal BC, and to the later 4th millennium, respectively. Stable isotopes suggest a diet based heavily on freshwater fish, and this is supported not only by ten thousands of identified fish remains, but also by a fish bone concentration nearby the skull of the man, which is interpreted as remain of a grave gift (possible fish soup). Of special interest are the baby’s stable isotope values. It shows that the mother’s diet was atypical (perhaps because she was non-local), and/or that dietary stress during pregnancy increased fractionation between the mother’s diet and her bloodstream.

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Утверждено к печати Ученым советом ИИМК РАНПрограммный комитет конференции: д. и. н.В.А.Лапшин (ИИМК РАН, сопредседатель) д

ABSTRACT Archaeological material was initially discovered in 1993, eroding from a small cliff on the north side of the peninsula of Asnæs near the town of Kalundborg in western Sjælland, Denmark. Ertebølle Excavations in 2007 exposed the Ertebøllwe cultural layer and obtained materials to describe the site and its contents before it was destroyed by the sea. The 22 m2 of careful excavations exposed a terrestrial midden deposit and the late Mesolithic cultural layer which had been partially preserved under a raised beach ridge. The flint tools consist primarily of projectile points, flake axes, some distally concave truncated blade knives, and a very few scrapers. There were large numbers of well-preserved faunal remains including bone fishhooks and preforms, seal bones, large bird bones, and an extraordinary amount of fish bone. A quantity of pottery was recovered in the excavations as well, including both pointed-bottom vessels and oval lamps in different sizes from the late Mesolithic and several examples of what are probably Early Neolithic ceramics. The rich occupation layer with its diverse artifactual content, including a fragment of a human jaw, documents a sizable residential settlement on the north coast of the Asnæs peninsula.

Утверждено к печати Ученым советом ИИМК РАНПрограммный комитет конференции: д. и. н.В.А.Лапшин (ИИМК РАН, сопредседатель) д

Утверждено к печати Ученым советом ИИМК РАНПрограммный комитет конференции: д. и. н.В.А.Лапшин (ИИМК РАН, сопредседатель) д

Archaeological material was initially discovered in 1993, eroding from a small cliff on the north side of the peninsula of Asnæs near the town of Kalundborg in western Sjælland, Denmark. Ertebølle Excavations in 2007 exposed the Ertebøllwe cultural layer and obtained materials to describe the site and its contents before it was destroyed by the sea. The 22 m2 of careful excavations exposed a terrestrial midden deposit and the late Mesolithic cultural layer which had been partially preserved under a raised beach ridge. The flint tools consist primarily of projectile points, flake axes, some distally concave truncated blade knives, and a very few scrapers. There were large numbers of well-preserved faunal remains including bone fishhooks and preforms, seal bones, large bird bones, and an extraordinary amount of fish bone. A quantity of pottery was recovered in the excavations as well, including both pointed-bottom vessels and oval lamps in different sizes from the late Mesolithic and several examples of what are probably Early Neolithic ceramics. The rich occupation layer with its diverse artifactual content, including a fragment of a human jaw, documents a sizable residential settlement on the north coast of the Asnæs peninsula.

The aim of this research is to examine the isotopic characterisation of archaeological fish species as it relates to freshwater, brackish and marine environments, trophic level and migration patterns, and to determine intraspecies variation within and between fish populations in different locations within central and northern Europe. Carbon and nitrogen stable isotope analysis was undertaken on collagen extracted from 72 fish bone samples from eight Mesolithic and Neolithic archaeological sites in this region. Thirty-six (50%) of the specimens analysed produced results with acceptable carbon to nitrogen atomic ratios (2·9–3·6). The fish remains encompassed a wide spectrum of freshwater, brackish and marine taxa (n = 12), which were reflected in the δ13C values (−24·5 to −7·8‰). The freshwater/brackish fish (pike, Esox lucius; perch, Perca fluviatilis; zander, Sander lucioperca) had δ13C values that ranged from −24·2 to −19·3‰, whereas the brackish/marine fish (spurdog, Squalus acanthias; flatfish, Pleuronectidae; codfish, Gadidae; garfish, Belone belone; mackerel, Scomber scombrus) ranged from −14·9 to −9·4‰. Salmonidae, an anadromous taxon, and eel (Anguilla anguilla), a catadromous species, had carbon isotope values consistent with marine origin, and no evidence of freshwater residency (−12·7 to −11·7‰). The δ15N values had a range of 6·2‰ (6·5–12·7‰) indicating that these fish were on average feeding at 1·7 trophic levels higher than their producers in these diverse aquatic environments. These results serve as an important ecological baseline for the future isotopic reconstruction of the diet of human populations dating to the late Mesolithic and early Neolithic of the region.

Recent excavations at two rockshelters (Olsteinhelleren and Sævarhelleren) on the Hardanger fjord in Western Norway have provided an unparalleled opportunity to examine the Mesolithic subsistence economy of this region. Thousands of fish remains (as well as numerous mammal and bird bones) have been analysed from these assemblages. Results show that the fishery was dominated by gadids, but labrids and salmonids were also important. Many other fish were present in small quantities, including the first specimen of sturgeon from the Stone Age of Norway. The transition to a more specialised fishery at the younger site, Olsteinhelleren, may be a reflection of a switch to the use of this locality as a logistic camp for the targeting of gadid fish.